(Spearman rank correlation r s = .333, P < .05) 

 for non-adjacent tissues is not statistically 

 - .133, P > .05). The ratio of A:B is plotted 

 demonstrates that the relative proportion of p 

 tentacles on adjacent regions increased during 

 period. Figure 3 shows the cumulative changes 

 number of polyps with sweepers on adjacent (A) 

 (B) regions at various times after initiation 

 These data suggest a cumulative decrease in th 

 on tissues not adjacent to colonies of _M. a nnu 

 clearly true for at least two colonies of M. c 

 sweeper tentacles on non-adjacent tissues disa 

 experiments were begun (4 and 6 weeks, respect 

 Necrotic wounds appeared on most colonies o 

 close to the colonies of M. cavernosa . Most ( 

 formed in intervals between observations, duri 

 number of M. cavernosa polyps with sweeper ten 

 the M. annularis also increased. Sweeper tent 

 cavernosa often were observed touching live M. 

 close to necrotic regions. 



. The 

 s i g n i f i 

 in fig 

 ol yps w 

 the ob 

 (media 

 and no 

 of the 

 e numbe 

 1 a r i s . 



avernos 



correl ati on 

 cant ( r s = 

 ure 2 and 

 i t h sweeper 

 servat i on 

 n ) in the 

 n-adjacent 

 experiment . 

 r of sweepers 



This was 

 a on which 



after the 



ppeared 

 ively) . 



f M. annularis 

 5 of 67 



n g w h i c 

 tacl es 

 acl es o 

 annul a 



wounds 

 h time the 

 adjacent to 

 f the M. 

 ris tissues 



DISCUSSION 



M 

 swee 

 with 

 numb 

 us ( 

 per 

 tent 



M 

 with 

 port 

 poss 

 the 

 i ncu 

 tent 



A 

 woun 

 Chor 

 cave 



cavernosa 



pert 

 ot he 

 er of 

 SLW) 

 polyp 

 acl es 

 oreov 

 swee 

 i ons 

 ibly 

 inter 

 rred 

 acl es 

 1 thou 

 ds in 

 nesky 

 rnosa 



thei 

 si mi 

 M. a 

 dama 

 For 

 grad 

 swee 

 at a 

 cave 



r rea 



1 ar t 

 nnu 1 a 



ge to 

 examp 

 ual 1 y 

 pert 

 grea 

 rnosa 



by M 

 et a^ 



and 



a nn 



entacl 

 r cora 



polyp 

 sugges 



and t 



may i 

 er , ou 

 per te 

 of col 

 may de 

 action 

 by the 



close 

 gh cau 



coral 

 , 1983 



we re 

 ch. W 

 o thes 

 ri s 



M. a n 

 le, if 



i nto 

 entacl 

 t e r d i 



sweep 

 u 1 a ri s 



app 

 es c 

 Is. 

 s ha 

 t th 

 he s 

 ncre 

 r da 

 ntac 

 onie 

 crea 



i nc 



pro 



to 

 ti on 

 -cor 



). i 



capa 

 heth 

 e ex 

 deve 

 nul a 



aren 

 1 ose 



Alt 

 ving 

 at , 

 i ze 

 ase 

 ta s 

 1 es 

 s no 

 se a 

 reas 

 duct 

 the 



sho 

 al i 

 tap 

 ble 

 er t 

 peri 

 1 opm 

 ri s 



1-. 1 

 exper 



979; s 

 imenta 



col 

 cont 

 es p 

 stan 

 er t 



and 

 ee C 

 1 in 



on i e 

 act , 

 ri or 

 ce t 

 enta 

 /or 

 horn 

 tera 



tly 



to 

 houg 



swe 

 i n a 

 of t 

 cl OS 

 ugge 

 ore 

 t i n 

 s th 

 es . 

 i on 

 si te 

 uld 

 nter 

 pear 

 of d 

 he c 

 ment 

 ent 

 by M 

 s of 



M. 



to 

 han 

 cl es 

 dige 

 esky 

 cti o 



can 

 the 

 h ou 

 eper 

 ddit 

 he a 

 e to 

 st t 

 xpan 

 vol v 

 e nu 



If 

 and/ 



of 

 be e 

 acti 

 s th 

 amag 

 ours 



s (i 

 of s 

 . ca 



i ncr 

 si te 

 r da 



ten 

 ion, 

 cros 



the 

 hat 

 si on 

 ed i 

 mbe r 

 so , 

 ore 

 comp 

 xerc 

 ons 

 at t 

 ing 

 e of 

 • e. , 

 weep 

 vern 



_M . annu 

 ca vernos 



ease the 



of comp 

 ta speci 

 tacl es , 



the num 

 pheres o 



site of 

 ei ther t 



of swee 

 n compet 



of expa 

 this m i g 

 x p a n s i o n 

 eti ti ve 

 i sed in 

 (see Bak 

 he sweep 

 tissues 



natural 



di gest i 

 er tenta 

 osa swee 

 1 a ri s an 



d i ge 

 the 



mig 

 sti o 

 , 19 

 ns s 



sti o 



1 cm 

 ht e 

 n by 

 83, 

 omet 



a may be 

 n by M. 



gap in 

 f f i ci ent 



M. a nnu 



for disc 

 i m e s may 



number 

 e t i t i v e 

 f i cal ly 

 observat 

 ber of s 

 n e x i s t i 



such en 

 he numbe 

 per tent 

 i t i v e in 

 nded swe 

 ht ref 1 e 



of a d d i 

 encounte 

 a s c r i b i n 

 , _et al . 

 er tenta 

 of M. a_n 



i nterac 

 on of M. 

 cl es by 

 p e r s ) is 

 d _M. ca v 



able to 

 annul a ri 

 these ex 

 ly deter 

 1 a ri s [s^ 

 u s s i o n o 



differ) 



of polyps with 

 encounters 

 reported the 

 ions by one of 

 weeper tentacles 

 ng sweeper 

 counters . 

 r of polyps 

 acl es on 

 teracti ons 

 epers close to 

 ct a "cost" 

 tional sweeper 

 rs . 



g causes for 

 , 1982; 

 cl es of M. 

 nularis within 



t i ons is 



cavernosa by 

 M . cavernosa 



less clear. 

 ernosa g row 



devel op 

 s . Similarly, 

 pe ri ment s , ^. 



close g rowth 

 ensu Ri chardson , 



f how natural 



65 



