is supported by nutrient recycling in 

 the marsh (Haines et al. 1975; Chalmers 

 et al. 1976). Therefore, based on the 

 information available, nutrient regener- 

 ation by microbenthos and macrobenthos 

 and tidal input of particulate organic 

 carbon (detritus) play an essential role 

 in coastal ecosystem development. 



LITERATURE CITED 



Bancroft, K., E.A. Paul, and W.J. Wiebe. 

 1976. The extraction and measure- 

 ment of adenosine triphosphate from 

 marine sediments. Limnol .Oceanogr. 

 21:473-480. 



Cammen, L.M. 1976a. Accumulation rate 

 and turnover time of organic carbon 

 in salt marsh sediment. Limnol. 

 Oceanogr. 20:1012-1015. 



Cammen, L.M. 1976b. Macroinvertebrates 

 colonization of Spartina marshes 

 artifically established on dredged 

 spoil. Estuarine Coastal Mar. Sci. 

 4:357-372. 



Chalmers, A.G. , E.B. Haines, and B.F. 

 Shen. 1976. Capacity of a Spartina 

 salt marsh to assimilate nitrogen 

 from secondary treated sewage. 

 Tech. Completion Rep. USDI/OWRT 

 Project A-057-GA, Environ. Resour. 

 Cent., Georgia Inst. Tech., Atlan- 

 ta. 



Christian, R.R., K. Bancroft, and W.J. 

 Wiebe. 1975. Distribution of 

 microbial adenosine triphosphate 

 in salt marsh sediment of Sapelo 



Island, Georgia. Soil Sci. 119: 



89-97. 

 Garbisch, E. W. , Jr., P. B. Waller, and 



R. J. McCallum. 1975. Salt marsh 



establishment and development. 



Tech. Memo. 52, U.S. Army Corps 



Engin. Coastal Engin. Res. Cent. , Ft. 



Belvoir, Virginia. 

 Haines, E. B., A. G. Chalmers, R. B. 



Hanson, and B. F. Sherr. 1975. 

 pools and fluxes in a 

 salt marsh. In M. L. 



Nitrogen 

 Georgia 

 Wiley, 

 Vol. 3. 

 Hanson, R. B 



ed. Estuarine research, 



and W. J. Wiebe. 1977. 

 Heterotrophic activity associated 

 with particulate size fractions in 

 a Spartina alterniflora salt-marsh 

 estuary, Sapelo Island, Georgia, 

 and the continental shelf 

 Mar. Biol. 42:321-330. 

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 W. B. Saunders Co., Phila- 



U.S.A 



waters. 

 Odum, E. P. 



ecology. 



delphia. 

 Stevenson, L. 



574 pp. 

 H., C. E. Millwood, and B. 

 H. Hebeler. 1972. Aerobic, hetero- 

 trophic bacterial populations in 

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 268-285 in R. R. Colwell and R. Y. 

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ACKNOWLEDGEMENTS 



Photographs of the Buttermilk Sound 

 Habitat Development Site were provided 

 by Dr. R.J. Reimold. 



52 



