Lutz, Triploicl Mutants in Oenothcra. ]()} 



"Several of the figures (figs. 37, 38, 39) show that the number 

 of chromosomes in the 0. LamarcMana hybrid is greater than in 

 (>. Iain. A count has not been made in the prophase, but from 

 numerous counts in the metaphases and anaphases, the sporophyte 

 number is found to be at least twenty and probably very near 

 that number (fig. 37). It will be remembered that the sporophyte 

 number in 0. lain is 14" 19 ). 



He did not here state the number of Lauiarckiana hybrids in 

 which he found at least 20 chromosomes, but in a paper published 

 in July, 1907 (8, pp. 10 11), in which the offspring of this cross 

 are again discussed, he said: 



"It is doubtless true that chromosome counts as ordinarly made 

 come from a very few individuals. In the case of the Lamarclriaua 

 hybrid I have examined material from two individuals and from 

 different flowers in each; these plants were grown in different 

 seasons. In every count the number of chromosomes in the somatic 

 tissues has been twenty or twenty-one, though I have been as yet 

 unable to determine with certainly between these two numbers." 



ml 



In the summary on page 20 he said: 



; 'The number of chromosomes is now found to be twenty or 

 twenty-one constantly in all the plants of the LamarcMana hybrid 

 examined, while the number in the pure 0. Lamarckiana, as well 

 as in 0. lata, is fourteen, the reduced number being seven. . . . Thus 

 the pure 0. Laniarckiaitct and the LamarcMana hybrid, which are 

 identical in external appearance, differ widely in the number of 

 their chromosomes." 



In again referring to this supposed culture of (). lata X 0. 

 Lamarckiana in a short paper (10) published in January, 1908, 

 he said: 



"0. lata from a cross was found to have 14 chromosomes as 

 sporophyte number, but quite unexpectedly one of the other plants 

 from what was believed to be pure seeds of 0. lata X 0. LamarcMana 

 was found to contain 20 chromosomes." 



Speaking of irregularities of chromosome distribution in the 

 heterotypic mitosis he said: 



"Such irregularities will probably also explain other variations 

 of one or two in the chromosome numbers, including the 20 instead 

 of 21 chromosomes in what I-have now shown to be an 0. lata X 0. 

 <ji(jas hybrid." 



On the next page in the same note: 



"I have also recently examined material from another plant in 

 the cross which was believed to be pure 0. lata X 0. Laii/aiT/,-f'ana, 



19) Notwithstanding my first statements in regard to chromosome numbers in 

 0. lata (see foot-note 11), it will be later shown that the specific number for this 

 form is 1,1. 



