Acknowledgments 



Many people have helped me toward the completion of this study in ways big and small. 

 Over the years I have undoubtedly forgotten the contributions of some of them, and I 

 apologize for this. Of those I have not forgotten, I want to thank the following people for 

 lending me specimens under their care: Pere Alberch, Walter Auffenberg, James Berrian, 

 Robert Bezy, Steven Busack, Joseph Collins, Ronald Crombie, Mark Dodero, Robert 

 Drewes, William Duellman, Anne Fetzer, George Foley, Harry Greene, L. Lee Grismer, 

 W. Ronald Heyer, J. Howard Hutchinson, Charles Meyers, Peter Meylan, Mark Norell, 

 Gregory Pregill, Jose Rosado, Albert Schwartz, Jens Vindum, Van Wallach, John Wright, 

 George Zug, Richard Zweifel, and especially Jay Savage and Richard Etheridge whose 

 collections provided the majority of the specimens examined in this study. 



I am also grateful to various teachers, friends, and colleagues who helped my ideas on 

 systematics and iguanine biology unfold through countless discussions: Troy Baird, Aaron 

 Bauer, Theodore Cohn, Michael Donoghue, Richard Estes, Richard Etheridge, Jacques 

 Gauthier, Eric Gold, David Good, George Gorman, Scott Lacour, Eric Lichtwardt, James 

 Melli, Sheldon Newberger, Mark Norell, Michael Novacek, David Wake, and Andre 

 Wyss. Linda Condon-Howe, Charles Crumly, Sanae and John Moorehead, Douglas 

 Preston, Doris Taylor, and the late Kenneth Miyata generously provided lodging while I 

 was visiting museums. Richard Estes, Richard Etheridge, Darrel Frost, Gregory Pregill, 

 David Wake, and Edward Warren, provided valuable comments on earlier versions of the 

 manuscript. David Cannatella and Rose Anne White gready assisted in the preparation of 

 camera-ready-copy. 



Finally, I want to give special thanks to Karen Sitton for providing emotional support 

 in her unique and charming way and to Richard Etheridge and Richard Estes for their 

 influence on both my academic and personal development. 



This study partially fulfilled the requirements of a Master's degree in Zoology at San 

 Diego State University, but was completed at the University of California, Berkeley. The 

 research and preparation of the manuscript were supported in part by a grants from the 

 Society of Sigma Xi, the San Diego State University Department of Zoology, the Theodore 

 Roosevelt Memorial Fund of the American Museum of Natural History, and the Graduate 

 Student Research Allocation Fund of the Department of Zoology, University of California 

 at Berkeley. 



XI 



