mary (and perhaps secondary) pro- 

 duction suggest that this is not an 

 important coupling, but we lack long- 

 term data from estuaries, and it is 

 not clear to me how short-term, rapid 

 recycling of nutrients is linked to 

 long-term inputs. The rivers have 

 been flowing for a long, long time, 

 and we know that most estuaries have 

 been filling in with sediment in 

 spite of a rising sea level. But, 

 except for highly enriched urban 

 areas, we do not know if they are 

 also becoming more eutrophic. Fresh- 

 water inputs may yet prove to play a 

 role in the long-term fertility of 

 estuaries . 



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