Broadhurst: Struthiopteris in North America 259 



related species first in Osmunda and later (following Hoffman*) in 

 Onoclea. Later Willdenow and Brown each published new generic 

 names for part of the genus under discussion. Willdenow, f in 

 1809, published the genus Lomaria, citing all of Onoclea of Swartz 

 except 0. sorbifolia as the basis of the name; and in the following 

 year Willdenow { listed under Lomaria most of Swartz's species 

 of Onoclea. ' In 1810 Robert Brown§ published Stegania, before 

 mentioned, for eight African members of this genus. In so doing 

 he separated Spicant from its relatives, just as Swartz had done 

 when he placed it under Blechnum, and its relatives under Osmunda 

 and later Onoclea. 



This confusion has continued even to the present, although two 

 authorities, Trevisan || and Underwood,! have shown independently 

 (1) that Osmunda Spicant L. has "first claim to Struthiopteris, 

 becoming Struthiopteris Spicant" and (2) that the correct generic 

 name for Osmunda Struthiopteris L. is not Struthiopteris but Mat- 

 teuccia, established by Todaro** in 1866. 



Trevisan attempted to divide the genus Struthiopteris, recog- 

 nizing Lomaria Willd. for part of it. In his key he describes 

 Struthiopteris as having median sori and ascribes marginal sori to 

 Lomaria. Trevisan discusses but three of the North American 

 species, Spicant, doodioides, and polypodioides, which he places 

 under Struthiopteris. A careful examination of the North Ameri- 

 can material in the New York Botanical Garden herbarium does 

 not justify this division. The term "median" is not applicable, 

 even relatively, to the indusium in any of this material ; the sterile 

 extension outside the indusium may be perceptibly wider in the 

 young leaves, but the conditions in the mature fronds do not 

 justify the use of the term. Nor do S. Spicant and 5. polypodioides 

 differ from any of the other species in the relative position of the 

 indusium. 



* Deutsch. Fl. 2: 12. 1795. 

 t Ges. Nat. Fr. Bed. Mag. 3: 160. 1809. 

 JSp. PI. 5: 289. 1810. 

 § Prod. Fl. Nov. Holl. 152. 1810. 

 ' || Atti 1st. Yen. III. 14: 556. 1869. 

 If Mem. Torrey Club 6: 257. 1899. 



** Giorn. Sci. Nat. 1: 208-254. 1866. Todaro also recognized the priority of 

 Struthiopteris Spicant for Osmunda Spicant L. 



