164 Boeshore — The Morphological Continuity of 



The calyx of Aphyllon is 5-toothed, the teeth equal to, or 

 longer than the tube. Glandular hairs are numerous and dis- 

 tributed over the entire outer surface of the calyx; shortly, these 

 consist of a short broad basal cell, then a long cylindric cell, 

 next to this 2 to 3 cells decreasing in size, and finally the top or 

 capitate part of several rounded cells. Stomata are numerous 

 and of the normal type. 



In Conopholis the calyx is orbicular, split in front, and toothed 

 at the tip. Hairs and stomata are negligible in this genus. 



Stamens. 



The stamens in parasitic Scrophulariaceae and Orobanchaceae 

 are 4, didynamous, in Gesneraceae, 5 to 4 to 2. 



As to anthers, a totally different relation holds between par- 

 asitic Scrophulariaceae and Orobanchaceae on the one hand as 

 compared with Gesneraceae on the other. In the former the 

 top of the filament is broadly inserted into the swollen back of 

 the anther, the lobes of which in the parasitic Scrophulariaceae 

 and Orobanchaceae are prolonged above the connections, but 

 in a striking manner are prolonged downward as two parallel 

 or divergent awns. These anther lobes in all except two gen- 

 era of Scrophulariaceae (Harveya, Hyobanche) and two genera 

 of Orobanchaceae (Campbellia, Aeginetia) are like each other, 

 that is, are equally paired anther lobes. But in Harveya one 

 anther lobe is large, normal, and polleniferous, and prolonged 

 below into a long horn; the other lobe is small, abortive, and 

 radiates back from the top of the filament. The latter anther 

 lobe in Hyobanche has been entirely absorbed, so that now one 

 fertile anther lobe dehisces by a single basilar pore. 



It is of interest to find that in Christisonia and Aeginetia of 

 the Orobanchaceae a similar structure exists, for, as in Hyo- 

 banche and Harveya, each bears a single fertile anther lobe. 



As to antherine structure, this exactly agrees in parasitic 

 Scrophulariaceae and Orobanchaceae and totally differs from 

 anything encountered in Gesneraceae. The structure and ap- 

 pearance of such genera as Melampyrum, Tozzia, Euphrasia, 

 Pedicularis, Buchnera, Gerardia, Bartsia, etc., absolutely resemble 

 those of Orobanchaceae, and this the writer would regard as 

 one of the most important points of contact between the two 

 families. 



