626 A MANUAL OF THE PENICILLIA 



2.0 to 2.5 cm. in 2 weeks, more or less zonate and lightly furrowed in a 

 radial pattern, with surface appearing more or less fibrous to floccose, 

 tearing easily, predominantly in buff to orange-pink shades, light sporing 

 in dull gray -green shades usually affecting the colony appearance only in 

 central areas; exudate lacking; odor evident, difficult to characterize; re- 

 verse variously colored in greenish gray to dark gray shades or at times 

 orange-red shades near salmon or apricot buff (Ridgway, PI. XIV) in a 

 broad marginal zone ; conidiophores arising as short perpendicular branches 

 from aerial hyphae or ropes of hyphae mostly 30 to 50/x by 2.5 to 3.0m sep- 

 tate, with walls comparatively heavy and colored in dull yellow-green 

 shades, smooth or slightly rough, sometimes arising from short cells sug- 

 gesting the foot-cells of Aspergillus (fig. 160F); penicilli typically biver- 

 ticillate and symmetrical as described and illustrated by Smith (fig. 160E) 

 but in our cultures commonly fractional, ranging from monoverticillate 

 through asymmetric structures with 2 or 3 metulae to occasional larger and 

 symmetrical penicilli, metulae 8 to 10/x by 1.8 to 2.2)u with walls colored 

 as the conidiophores; sterigmata mostly in verticils of 4 to 6, about 8 to 10/i 

 by 1.5 to 2.0m, lanceolate with conidium-bearing tubes characteristically 

 tapered; conidia ovate to elliptical, up to 3.0 to 3.5m by 2.0 to 2.5m, thin- 

 walled, smooth. 



Colonies on steep agar somewhat larger than on Czapek but similar in 

 pattern and texture; reverse in shades listed above; penicilli averaging 

 slightly larger than above but otherwise similar in pattern. 



Colonies on malt agar spreading broadly (fig. 160D), up to 6 cm. in 2 

 weeks, seldom furrowed, consisting of a fairly loose, floccose to funiculose 

 mycehal felt, up to 1 mm. deep, lightly sporulating throughout, in light 

 gray-green shades near mineral gray (R., PI. XL VII); exudate lacking; 

 odor indefinite; reverse in dirty yellow-orange to almost black shades; 

 conidial structures as described on Czapek; metulae and sterigmata con- 

 sistently longer and thinner, usually closely parallel; conidia strongly el- 

 liptical to almost cyhndrical, about 2.5 to 3.0m b}^ 1.5 to 2.0m. 



The type strain, received from the National Collection of Type Cultures 

 as George Smith's culture No. 91, is maintained in our Collection as NRRL 

 2096. The same culture received from the Centraalbureau duplicates this 

 completely. 



This unique species has apparently been isolated only once. Its funicu- 

 lose habit described and illustrated by Smith and observed in all of our 

 cultures, unmistakably placed it in the Penicillium fimiculosum series. It 

 differs markedly from other members of the series in the production of 

 unusually short, conspicuously septate conidiophores with strongly pig- 

 mented walls arising from aerial hyphae usually colored in somewhat 

 lighter shades. The type culture has apparently changed markedly during 

 the period of laboratory cultivation. Smith originally figured it as pro- 



