REVISION OF ENDOGONEAE. 303 



The peculiar characters of this species illustrate the culmination of 

 the tendency toward a definite grouping of the spores within the gleba, 

 which is present to a less marked degree in E. tuberculosa and E. 

 fuegiana. The sexual nature of the spore-origin is unquestionable 

 from the two distinct origins are present in all spores. The alterna- 

 tive that they may be intercalary and represent a lateral bulging, so 

 to speak, in the continuity of the hypha, is an explanation which is 

 rendered quite improbable by our knowledge of spore-formation in all 

 the chlamydosporic types. The conjugation is evidently somewhat 

 peculiar, as is evidenced by the often remote origins, and it is to be 

 regretted that, owing to the fact that the whole spore-mass is hardly 

 distinguishable from a slightly coherent mass of earth, the younger 

 stages are not likely to be found, unless by accident. 



Endogone tuberculosa Lloyd. 

 (Figs. 11-16.) 

 Lloyd (1918), p. 799; fig. 1239. 



This species has been described and its gross appearance well illus- 

 trated by Lloyd, to whom the writer is indebted for a small portion of 

 the type material on which the following notes are based. It was col- 

 lected in New South Wales by Mr J. B. Cleland, who states that it 

 was found just at the surface of the ground, apparently partly buried 

 in it, if one may judge by the coating of earth which completely 

 envelopes it. Its gross characters are peculiar from the fact that the 

 gleba is not a continuous and undifferentiated spore-mass, but is in a 

 sense compound. 



The sporogenous area, which is only visible in sections. Figure 11, is 

 very irregular in outline, pushing indeterminate lobes or extensions 

 outward into the surrounding covering of earth, which thus varies 

 greatly in thickness, and appears to be held together by a scanty 

 penetrating mycelium. It is possible, after slightly moistening the 

 cut surface, to determine that the golden yellow spores are arranged in 

 rounded masses of variable size and shape, or are associated in larger 

 somewhat less definite areas. In either case they are often, though 

 not always, separated by intruding layers of the earthy matrix, the 

 presence of which is indicated by its darker color, and which may be 

 even more intimately incorporated in the general mass, although none 

 appears to occur within the individual spore-groups. 



In these spore-groups, or areas, the more clearly defined of which 



