1910] Brandt: Notes on Trillium 45 



will be noticed that the leaves greatly surpass the sepals and are folded 

 several layers thick around the flower-bud and the anthers at this 

 time practically fill the space in the center of it. Vascular bundles 

 are plainly distinguishable in the young stem. &., is the second normal 

 bud, which is shown entire. As in ho, the young foliage leaves com- 

 pletely enwrap the flower bud. The size and shape of the "axillary 

 sheath" is illustrated by x^. Sheath s^, as will be noted, is consider- 

 ably shorter than bud 63, so bud h^ would naturally be much smaller 

 than h.,. Only the position that bud h^ would occupy can be indicated. 

 Bud h\ will be noted under a very short, young sheath. This bud if 

 shriveled up would be a trifle smaller than h^, and perhaps a little less 

 advanced. It is not a bud for next season, but for the season following, 

 when it will fail of development in the same way as &i. b'2 is barely 

 distinguishable, and at a the apex of the rootstock is covered by a 

 few young sheaths which are merely arched and have no buds, only 

 "axillary sheaths," in their axils. 



It is important to note the abortion of h^ and the possible abortion 

 of h\. Apparently all the buds from fe. and inward were laid down 

 in the preceding active season, but h\ and 6'._, were arrested in devel- 

 opment so that they could not become shoots during the next season. 

 There is apparently a critical period during which further develop- 

 ment in the younger buds is temporarily suspended. Examination 

 of a crown late in May seemed to show that there is a selective elimi- 

 nation relative to the amount of differentiation the various buds ex- 

 hibit. Thus, as a general rule, the bud which fails during this critical 

 period to meet the requirements of sufficient growth dies and shrivels 

 up later on. In very strong plants of T. sessile var. giganteum, those 

 which send up four or more shoots in a season, the largest arrested 

 bud develops into a shoot, usually smaller than the one next within. 

 In very large plants this outer shoot may show increase in number 

 of parts or may even become fasciated. A report upon numerous 

 teratological forms peculiar to T. sessile var. giganteum is in progress 

 of preparation. 



The "crown" of T. ovatum is very similar in structure to that of 

 T. sessile var. giganteum. Usually not over two buds are formed each 

 year instead of two or more, as in the latter species. Tlie first of these 

 fails to develop. When the plant is given an unusual amount of light 

 both buds develop into strong, normal shoots. Because of its habit of 

 forming two buds a year, one may often find T. ovatum in partially 

 cleared forest areas regularly sending up two shoots from one root- 



