308 University of California PuhUcations in Botany [Vol. 10 



Marchantia Setchellii sp. nov. 

 Plate 83 



Dioicous. Medium size; dark glaucous green in color; densely 

 caespitose. Fronds dicliotomous, ligulate, equal breadth, cross-section 

 plano-convex, 20 to 25 cells thick at the middle, cortical cells two 

 laj^ers, dark-colored, similar in size to the inner, margin of fronds 

 often recurved, marginal cells smaller than others. Scales light 

 purple, oval, rotundate or obtuse, distinctly dentate, teeth short; 

 appendages deep purple, lanceolate-acuminate or ovate-acuminate, 

 dentate, teeth often long and hamate; cells oblong-quadrate. 



Stomata numerous, round or oblong, three tiers of cells high, 8 

 marginal cells, inner quadrate. 



Cupules shallow, dilated, fringe dentate, teeth 1 to 3 cells long. 



Dimensions. — Fronds 1 to l^/o inch long; 2 to 3 mm. wide; .5 mm. 

 thick at the middle, margin .1 mm. thick, marginal cells .01 to .02 

 mm. ; median scales .4 mm. X .45 mm., .35 mm. X .3 mm., cells of 

 same .05 mm. X .02 mm. ; appendages .6 mm. long X 2 mm. broad, 

 cells of same .075 mm. X .05 mm. ; teeth of fringe of cupule .1 mm. 

 long ; stomata .6 mm. diam. 



Habitat. — Slopes near Waimangu, North Island, W. A. Setchell, 

 1904, no. 160. Type, Herb. Univ. Calif., no. 213707. 



Observations. — Although the specimens received from Setchell are 

 all sterile, I have no hesitation in considering this a new species. 

 Evans, in The American Species of Marchantia (Trans. Conn. A. & S., 

 vol. 21, 1917), notes the specific value of the appendages; in this 

 species they are very distinct from any of those of any other species 

 recorded from New Zealand. The onlv one that has somewhat similar 

 ones is 31. papillata Raddi, a South American species, although 31. 

 platycnemos Schwaegr. from the Falkland Islands has been doubtfully 

 referred to this species. 



I have not had the opportunity of examining any specimens of 

 31. loapiUata except those of 31. subandina Spruce, which has been 

 reduced by Evans to a synonym of 31. papillata. 



31. subandina Spruce is similar in size, but the lobes are more 

 erect, not widely spreading, and in some forms f astigiate ; in a cross- 

 section they are 10 to 15 cells thick at the middle, which gradually 

 become thinner in the lamina. In the examination of 31. subandina 

 I noticed a singular feature. Many of the cross-sections showed, on 

 the ventral side, the bulging out of a semicircular band of 5 cells X 5. 

 This feature disappears on the younger fronds. 



In 31. Setchellii the fronds are 20 to 25 cells thick at the middle 

 and abruptly become thinner winged. 



