Superfamily ICHNEUMONOIDEA 479 



SUBTRIBE SPHECOPHAGINA 



This subtribe includes the Holarctic genus Sphecophaga and two other genera which are 

 restricted to the Old World. The hosts are pupae of Vespinae or Polistinae. 



Genus SPHECOPHAGA Westwood 



Sphecophaga Westwood, 1840. Introduct. Mod. Classif. Ins., v. 2, Synopsis Genera Brit. 

 Ins., p. 57. 



Type-species: A)io)naloh cespanoii Curtis. Monotypic and orig. desig. 

 Chrijono))iOH Desvignes, 1856. Cat. Brit. Ichn., p. 47. 



Type-species: Anoitialoii respanitu Curtis. Monotypic. 

 Cacotro'pa Foerster, 1868. Naturh. Ver. Rheinlande, Verh. 25: 208. 



Type-species: Cacotropa sericea Thomson. By subsequent monotypy from inclusion 

 by Thomson, 1888. The type-species is regarded as a synonym of S. vespariDn 

 I'espariDii (Curtis). 



There is a single species. 



vesparum burra (Cresson). N. H. w. to B. C, s. to S. C, Ariz., and Calif. Host: Vespula 



aroiaria (F.), V. )iiaci(lata (L.), V. )iiaculifro)ts (Buyss.). S. vesparum vesparuin occurs 

 in Europe. Both diapausing and nondiapausing larvae may develop on the same host 

 pupa presumably from eggs deposited by a single v. burra female. Only females develop 

 from nondiapausing larvae, and they usually differ somewhat in coloration from females 

 which develop from diapausing larvae. Nondiapausing larvae spin a delicate, white 

 cocoon, while diapausing larvae spin a tough, brown one. 



Eucero.s burriis Cresson, 1869. Canad. Ent. 1: 104. 9. 



Sphecophagml'V praedator Zabriskie, 1894. N. Y. Ent. Soc, Jour. 2: 84. 6, 9. 



Biology: Couper, 1870. Canad. Ent. 2: 52-53. -Zabriskie, 1894. N. Y. Ent. Soc, Jour. 2: 81, 86. 

 — Cushman, 1933. Ent. Soc. Wash., Proc. 35: 10-11. — Schmieder, 1939. Ent. News 50: 

 92-97, 125-131. -Leech, 1954. Pan-Pacific Ent. 30: 80. 



Unplaced taxa of CRYPTINI 



Cryptm niiindiis Provancher, 1874. Nat. Canad. 6 (6): 177 (key); 6 (7): 203. "S"= 6, 9. Que. 

 Barron (1975) said that the lectotype (no. 245 in second collection of Provancher) keyed 

 to Cryptus ruralis Pratt, a species not known from Quebec. Townes' (1944) placement of 

 niiDidiis as a synonym of C. persiviilis Cresson was based upon specimen no. 189 in 

 Provancher's first collection (personal commun.. Dr. H. K. Townes, 1975). 



Taxonomy: Townes, 1944. Amer. Ent. Soc, Mem. 11: 280. —Barron, 1975. Nat. Canad. 102: 

 511-512. 



Subfamily ACAENITINAE 



Revision: Cushman and Rohwer, 1920. U. S. Natl. Mus., Proc 57: 503-523. —Townes and 

 Townes, 1960. U. S. Natl. Mus. Bui. 216 (pt. 2): 544-579. 



Ta.xonomy: Townes, 1971. Amer. Ent. Inst., Mem. 17: 159-179 (genera of world). 



Tribe COLEOCENTRINI 



This Holarctic and Oriental tribe includes six genera, only two of which have North American 

 species. 



Genus COLEOCENTRUS Gravenhorst 



ColeocentniK Gravenhorst, 1829. Ichn. Europaea, v. 3, p. 437. 



Type-species: IclDienniou excitator Poda. Desig. by Westwood, 1840. 

 Macrocoleus Desvignes, 1850. Ent. Soc. London, Trans, (n. s.) 1: 13. 



Type-species: Macnifi longiventris Gravenhorst. Desig. by Viereck, 1914. The 

 type-species is regarded as a synonym of C. excitator (Poda). 



This is a moderately large Holarctic and Oriental genus. 



