NO. 1 hartman: goniadidae, glyceridae, nephtyidae 45 



gnaths are designated jaws, whereas the proximal proboscidial organs 

 are called paragnaths. The illustration of the terminal aperture, show- 

 ing macrognaths and micrognaths, is likewise reversed, so that the ven- 

 tral macrognaths are shown dorsal, and the dorsal arc of micrognaths 

 is shown below. Levinsen's description of the arrangement of the parts, 

 however, resembles that in other species of the genus. A translation 

 follows. 2 "The ventral [dorsal] paragnaths form a fairly broad, longi- 

 tudinal belt which is separated from the other by a narrow median 

 space [area I, textfig. 2]. In this space one finds a row of extremely 

 small pieces [proboscidial organs of area I] ; each longitudinal band is 

 again composed of series of transverse, or more correctly slightly slant- 

 ing rows of belts of which each is composed of 5 [though actually 6 

 are shown] pieces with a formula 1.1.2.1. Commonly these paragnaths 

 could be described as narrow, compressed bodies which are fastened by 

 a broad, basal portion and raised freely from the upper surface of the 

 proboscis. The innermost piece in each row [proboscidial organ II-2] 

 is hooklike and resembles much a hook of a Taenia. The hook is directed 

 diagonally backward. The next piece [II-3] has a diagonally cut-off 

 point and on the outermost the point is forked. Between each 2 hooks in 

 the same longitudinal row there lies yet a small narrow piece [II-l] 

 .... The dorsal paragnaths [ventral proboscidial organs] are on each 

 side separated from the ventral [dorsal] ones by a considerable space. 

 They are tongue-shaped, fairly short pieces, arranged in 4 widely sep- 

 arated, single, longitudinal rows [includes proboscidial organs of areas 

 IV and V]. The paragnaths in the 2 middle rows [V] are larger and 

 more widely separated from each other than in the 2 outer [IV]." The 

 details of the individual pieces are not shown. Thus, the pores which 

 are almost certainly present in most of the organs are neither shown nor 

 mentioned ; the number of processes, if any, on proboscidial organ IV 

 (see textfig. 2) is not disclosed. A more precise description of these 

 organs is lacking for G. nordmanni (Malmgren). 



These organs not only occupy well defined areas, but have char- 

 acteristic shapes. They are here described in detail since they are found 

 to be specific for all species investigated. For ready reference, the sur- 

 face of the proboscis is divided into areas, numbered I to VI. These 

 areas are most clearly distinguished in cross section of the proboscis 

 (see textfig. 2). Area I is mid-dorsal; it may have one to several rows 



2 I am indebted to Mr. Anker Petersen of the Allan Hancock Foundation 

 staff for a translation of this account from the Danish into English. 



