No. 5 meserve: monogenetic trematodes 33 



Oken, 1815 (P. hippoglossij. Phylline was shown by Odhner (1905) 

 to be a synonym of Tristomum maculatum Rudolphi, 1819 (which 

 in turn is now considered a synonym of Capsala). The type of 

 Epibdella is E. hippoglossi (Mueller, 1776) which possesses pseudo- 

 suckers. Odhner (1905) elevated the subgenus Benedenia to generic 

 rank, resulting in two genera, Epibdella with pseudosuckers and 

 Benedenia with true suckers. Johnston (1929) points out that Epib- 

 della should be submerged as a synonym of Entobdella. 

 Johnston (1929) says: 



In 1817 Cuvler, in his "Le Regne Animal," vol. 4, erected the genus Tristoma, 

 describing and figuring one species, T. coccineum. Next year Lamarck (1818, 295) 

 gave a summary of Phylline, mentioned the synonymy of P. hippoglossi, and stated 

 his belief that the parasites were related to Poly stoma instead of Annelids (leeches), 

 where they had been allotted. He also referred to Blainville's manuscript name, 

 Entobdella, for the genus, but retained Oken's Phylline. The reference was quoted 

 erroneously by Braun (1889) as appearing in Lamarck's vol. 1, p. 444, and subse- 

 quently (1890, 518) he indicated the genus with the date 1815 (when vol. 1 

 appeared) as a synonym of Epibdella. Stiles and Hassall (1908. 251) credit Entob- 

 della to Audouin 1828, whereas Agassiz (1845) and Scudder (1884), in their 

 respective Nomenclatores Zoologici, attribute it to Blainville, but without mentioning 

 a date. Sherborn, in his Index Animalium, gives the correct date (1818) for Entob- 

 della (Blainville MS.) Lamarck. It was not mentioned by Rudolphi (1819), and 

 has remained practically unrecognized since. 



Yamaguti (1937) uses the genus name Epibdella with two sub- 

 genera, Epibdella and Benedenia, and classifies it in the family Cap~ 

 salidae. He does not explain his adoption of these names but evi- 

 dently does not follow either Odhner or Johnston. The subgenus 

 Epibdella seems to possess anterior suckers and the subgenus Bene- 

 denia seems to possess anterior suckers together with the folds of 

 "an anterior adhesive organ," thus corresponding to the genera 

 Benedenia and Pseudobenedenia, respectively, of Johnston (1931). 



Johnston (1931) in a revision of the family Capsalidae Baird, 

 1853 {Tristoviidae Taschenberg, 1879) names the subfamily Cap- 

 s alinae ]o\in?,lon, 1931 {T ristominae MonlxcQWi, 1903) with Capsala 

 (Tristoma) as the type genus characterized by numerous radii in 

 the posterior haptor; the subfamily Nitzschinae Johnston, 1931 with 

 Nitzschia von Baer as the type genus characterized by numerous 

 testes and a median genital pore; Ancyrocotylinae Monticelli, 1902; 

 Benedeniinae Johnston, 1931. Johnston (1931) gives the following 

 diagnosis of the subfamily Benedeniinae: 





