34 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 2 



Capsalidae with, large stalked posterior suckers, provided with typically, three 

 pairs of hooks, the second pair largest, the third smallest (or perhaps absent), but 

 without septa dividing it into loculi; either with anterior glandular organs (Entob- 

 della) or with anterior suckers (Benedenxa), or with both (Pseudobenedenia) ; with 

 two testes lying side by side. 



I include the following subfamilies under the family Capsalidae: 

 Capsalinae Johnston, 1931, Nitzschinae Johnston, 1931, and Bene- 

 deniinae ]o\v!\%ion, 1931. 



The genera of the subfamily Benedeniinae may be distinguished 

 as follows: 



1 (2) Lateral intestinal branches absent 



Ancyrocotyle Parona and Perugia, 1903 



2 (1) Lateral intestinal branches present 3 



3 (4) True anterior suckers absent; anterior 



adhesive organs present 



Entobdella Blainville, 1818* 



4 (3) True anterior suckers present 5 



5 (6) Anterior adhesive organs present 



Pseudobenedenia Johnston, 1931 



6 (5) Anterior adhesive organs absent 



Benedenia Odhner, 1905 



According to Johnston (1931, p. 95) Ancyrocotylinae should be 

 restricted to include Ancyrocotyle with a single preovarian testis 

 and the absence of lateral branches to the intestinal crura. Price 

 (1934), however, places Ancyrocotyle in the subfamily Benedeniinae. 

 It was suggested by him that, due to the fact that the material of 

 Ancyrocotyle vallei was in poor condition, there might have been 

 an error in interpreting the preovarian testis. A. bartschi Price, 1934 

 has two testes posterior to the ovary. 



In this paper, the genus name Benedenia is accepted for those 

 forms with true anterior suckers. The following species of Benedenia 

 have been described: 



1. B. derzhavini (Layman, 1930) n. comb, on Sebastodes schle- 

 gelii. 



*Lamarck (1818) in referring to Phylline Oken states: "Ce genre est etabli 

 par M. Ochen, sous le nom que nous lui conservons, et neanmons M. de Blainville, 

 que I'avait deja reconnu, lui assigna celui de Entobdella, dans ses manuscrits." 



