91 



accompanied by a division of the cell wall (PI. XII, Fig. 101). 

 Associated with the archegonia are short glandulai' hairs or 

 paraphyses which usually consist of one short basal cell and a 

 longer terminal one. The dehiscence of the archegonium was 

 not studied in the living state, but opening archegonia were 

 frequently met with, and in a few cases the entrance of the 

 spermatozoids was demonstrated. The fertilization was not 

 studied in detail, but the entrance of the spermatozoid into 

 the egg and the penetration of the egg-nucleus seems to 

 closely correspond to what has been studied and observed in 

 other cases. 



Of the other species examined, G. pohjiwdioides differs from 

 G. pectinata in the character of the archegonium more than do 

 the other more nearly related species. In the former the arche- 

 gonium neck is very strongly bent forward and is longer and 

 contains more cells. The whole archegonium is larger than in 

 the other species. In this species also no archegonium was seen 

 in which the neck-canal -cell w^as divided by a wall. Only a 

 small number of archegonia were studied, however, and it is 

 quite possible that such division may sometimes occur. 



The archegonia in both G. ^/Mo^oma (PL XIII, Figs. 104—106) 

 and G. laevigata (PI. XIII, Figs. 109 — 110) closely resemble G. 

 pectinata both in form and size, and as in that species there are 

 usually two distinct neck-canal-cells. The forward curve of the 

 neck in all the species of Glelchenia is an interesting contrast to the 

 backward bending of that of the Poly pod iaceae. Probably in 

 both cases the position facilitates the entrance of spermatozoids. 



The Embryo. 



Only a small number of the embryos was secured and it 

 was not possible to follow as fully as could have been wished 

 all of the earlier divisions. Rauwenhoff figures sections of tw^o 

 young embryos of G. dlcarpa, but he did not follow in detail 

 the divisions beyond the earliest stages. The first division seems 

 to correspond to that of the Polypodiaceae, the first or basal 

 wall being vertical and coinciding approximately with the axis 



