77 



each of the four original neck-cells often dividing only once, 

 so that there may l:>e but two cells in each of the four rows. 

 More often there is a second division in some of the cells so that 

 each row consists of three cells (PI. VIII, Fig. 28). The nucleus of 

 the broad neck-canal-cell probably always divides, but in the few^ 

 sections where the two nuclei could be seen (PI. VIII, Figs. 26 — 28) 

 there was no indication of a wall between them, but as has 

 sometimes been observed in other Marattiaceae it is by no 

 means unlikely, however, that such a division may occur also 

 in Kaulfussia. The ventral canal-cell is conspicuous and equals in 

 breadth the sister-cell, the egg (PI. VIII, Fig. 27). The large venti-al 

 canal-cell in the Marattiaceae is in marked contrast to the 

 very inconspicuous one found in the Ophioglossaceae where it 

 is usually very difficult to demonstrate. In the peripheral 

 portion of all the axial cells of the arcJiegonium there are 

 many small starchgranules. Jonkman figures similar starch 

 grains in both Marattia and Angiojjteris. Mantle-cells like those 

 surrounding the antheridium may be formed, but they are not 

 so conspicuous and do not surround the whole of the arche- 

 gonium venter. In this respect there is also a close Ptgreement 

 between the Marattiaceae and Ophioglossum. 



The Embryo. 



Only two young embryos were secured so that it was im- 

 possible to follow in detail the development of the young 

 sporophyte. The first or basal division wall, as in other 

 investigated Marattiaceae, is transverse, and to judge from 

 a comparison with the embryos of Anglopteris, of which a 

 number of preparations were made, all of the organs ot the 

 young sporophyte except the foot are of epibasal origin, the 

 entire hypobasal half of the embryo forming the foot. In this 

 respect the Marattiaceae resemble Isoetes (see Campbell, Con- 

 tributions to the life-history of Isoetes, Ann. of Bot. V, 1891), 

 and Botrychiuin, where the whole of the epibasal half of the 

 young embryo also becomes the foot (Jeffrey, E. C, Gameto- 

 phyte of Botrychium Virginiamuii, Proc. Canadian Institute, V, 



