76 



fully developed when there seems to be a contriiction of the 

 Ducleus which becomes narrower and almost homogeneous. The 

 nucleus occupies only the large posterior coil of the spermatozoid, 

 all of the forward portion, about two small coils, being formed 

 of the blepharoplast with probably a certain amount of other 

 cytoplasmic matter. The free spermatozoids were not seen as 

 the material was collected just as I was on the point of leaving 

 Buitenzorg and it was necessary to preserve it at once. 



While the series of preparation of the developing spermatozoids 

 was not entirely complete it was clear that the development 

 follows very closely thtit of Opiiloylossum. 



The A r c h e g n i u m. 



The archegouia are formed as a rule only after the antheridia 

 have ceased to develop. They closely resemble those of other 

 Marattiaceae but are decidedly larger, and in this respect Kaul- 

 fussia is again more like Opluoglossum. (Lang, On the prothal- 

 lium of Opluoglofi.niiii pendulum and llelnimtho.stachys Zei/Ianicd, 

 Ann. of Bot., XVI, 1902; Campbell, loc. cit.) As in the case 

 of the antheridium the archegonia are formed acropetally. As 

 in all species of Marattiaceae the neck of the archegonium is 

 very short and hardly projects at all above the surface of the 

 prothallium. The first division in the mother-cell is like that 

 in the young antheridium and results in the separation of an 

 outer or cover-cell from the inner one (PI. VIII, Fig. 23). The 

 latter again divides by a periclinal wall into a ceutral cell, 

 from which later arise the egg-cell and canal-cells, and a 1)asal 

 cell {/)) which is probably always present, but is less conspicuous 

 than is the case with many other ferns. Like tlie mother-cell 

 of the antheridium there is a good deal of variation in the 

 width of the young archegonium. Some of the narrower types 

 (PI. VIII, Fig. 25) recall the archegonium of Anthoceros and suggest 

 the possible derivation of this type of archegonium from one like 

 that found in the Anthocerotaceae. The further development 

 of the archegonium corresponds almost exactly to that already 

 described in various Marattiaceae. The neck remains very short. 



