73 



and figures septate rhizoids in Danaea shnplicifolia and assumed 

 that they were really multicellular. He does not appear, how- 

 ever, to have made a critical examination of the nuclei, and 

 it is not unlikely that these rhizoids also are not really mul- 

 ticellular. 



The apex of the prothallium is much like that of other ferns. 

 The apical meristem lies at the bottom of the sinus and a 

 horizontal section (PI. VII, Fig. 4) shows the usual row of marginal 

 cells one of which is generally larger than the other and may 

 represent a single apical cell, but this is difficult to decide as 

 there is no doubt that all of the marginal cells act to some 

 extent as initial cells for the growth of the thallus, and in 

 longitudinal sections appear very much alike. As in other ferns 

 the longitudinal section appears nearly semi-circnlar, the cells 

 having in fact the form of half a disc. The principal segments 

 are cut off from the internal face, bnt lateral segments adding 

 to the row of marginal cells are also formed from time to time. 



In the interior cells of the older parts of the gametophyte 

 there occurs an endophytic fungus very much like that found in 

 the prothallium of the Ophioglossaceae (Pl.VII, Fig.lO). I have also 

 found this in Angiopteris and in Marattia Douglnsii, and it is 

 quite likely that it occurs largely in all Marattiaceae. As we 

 shall see later a similar endophyte occurs in many, if not all, 

 species of Gleiclienia. 



The Autheridium. 



The antheridia in Kaulfassia are developed upon the younger 

 prothallia before the archegonia appear. They seem to be con- 

 fined strictly to the underside of the midrib near the apex, 

 occupying relatively exactly the same position that the arche- 

 gonia do in the older prothallium. None were seen upon the 

 wings of the prothallium, nor were they in any cases found 

 upon the u])pei' side, as is by no means uncommon in Marattia 

 and Anyiopteris. They are much larger than those of any of 

 the other Marattiaceae that hiive been studied, and in this 

 respect as well as in the larger spermatozoids they resemble 



