164 



more or less conspicuous reticulum, but the nucleolus has ap- 

 parently disappeared, as it usually does at this stage in other 

 forms that have been studied. The further development of the 

 spermatozoid corresponds very closely w^ith that of Equisetnm 

 (see Campbell — Mosses & Ferns, 2d edit., p. 448 — Belajaff. 

 tlber die Cilienbildner in den Spermatogenen Zellen. Ber. der 

 deutsch. hot. Gesells., XVI: 140, 1898). One of the blepharo- 

 plasts in the primary spermatocyte goes with each daughter 

 cell (the definitive spermatocyte) and can be seen as a distinct 

 rounded body near the nucleus (Fig. 87 a). In some cases what 

 appears to be the blepharoplast lies in a depression at the 

 periphery of the nucleus and looks very much like a nucleolus. 

 Before the nucleus undergoes any marked change the ble- 

 pharoplast begins to elongate (Figs. 80—88) and assumes the 

 form of a pointed, slender, strongly stained body lying close 

 to the nucleus. This is really somewhat ribbon-shaped, and 

 more pointed at one end than the other. It is usually some- 

 what curved even in its earliest stages, and the transverse 

 section (Fig. 80^) appears crescent shaped. The nucleus next 

 elongates slightly and the reticulate structure becomes very 

 conspicuous (Figs. 90 — 91) with large strongly stained granules 

 which are probably composed of several more or less coherent 

 chromosomes, as the number of these is very much less than 

 the chromosomes in the nuclear plate of the dividing nucleus 

 of the primary spermatocyte. The blepharoplast continues to 

 grow, and at this stage (Figs. 91 — 92) is already strongly 

 curved, and in favorable cases the young cilia can be detec- 

 ted — but none of the writer's preparations showed this nearly so 

 plainly as Balajefe's figures for Equisetnm and Gymnogramme. 

 There is no doubt, however, that the cilia arise in much the 

 same way as he describes for those forms. The nucleus next 

 becomes indented on one side and assumes a crescent shape, 

 which elongates, becoming also more or less flattened. One 

 end becomes narrower and sharply pointed, the other remaining 

 thicker and rounded. The thick reticulum stains now with 

 great intensity, and shows a tendency to coalesce", which in 



