87 



Fig. 44 PI. IX shows two nuclei in the antipodal cell ')• 

 Fig. 46 PL IX shows the only instance in which I have seen 

 three well developped antipodes. The antipodal cell can occas- 

 ionally reach enormous size (fig. 42 PL IX) which may attain 

 to one third of the size of the embryosac. 



Consequently the eggapparatus is formed in a perfectly normal 

 way, and as it is formed at the organically upper top of the 

 embryosac, we have what v. Tieghem calls acrogamy. Van 

 TiEGHEM distinguishes two kinds of embryosacs viz. acrogamous, 

 and basigamous ones according to the positions occupied by the 

 eggapparatus at the originally upper or lower pole of the sac. 



Our figure 49 shows that in a normally acrogamous embryosac, 

 basigamy or at least almost basigamy may occur by the 

 eggapparatus attaching itself to the lateral wall of the em- 

 bryosac. 



At the moment when the two polar nuclei touch previous 

 to fusing (fig. 37, 39 PL VIII, fig. 45 PL IX) one would expect 

 especially after Nawashin's investigation to see the pollentube 

 entering the embryosac. Nawashin whose most mp.gnificent 

 preparations I had the good fortune to see proved without the 

 shadow of a doubt; that the endospermnucleus of Fritillaria 

 results from the fusion of three nuclei viz. the two polar nuclei 

 and one of the two generative, wormshaped nuclei, the remaining 

 one of the generative nuclei copulating with the eggnucleus ^). 



Notwithstanding most numerous trials on a material as large and 

 containing as many different stages as certainly nobody had previously 

 at his disposition I have never seen a pollentube enter the embryosac, 

 nor did I ever See a pollengrain attached ') to the stigmata , allt hough 

 these latter are so small that no pollengrahi , if present , could 

 possibly escape detection. 



1) This antipodal cell ressembies very much the one of Gonyanthes Candida 

 (Burmanniaceae) see Treub, Annales d. Jard, Bot. de Buitenzorg. Vol, III 1883 

 fig. 1 and 2 PI. XVIII. 



2) Sergius Naivaschin. Resultate einer Revision der Befriichtungsvorgange bei 

 Lilium Martagon und Fritillaria tenella. Bulletin de TAcaddmie Imp^riale des 

 sciences de St. P^tersbourg. 1898. Novembre. T. IX n" 4. 



3) Except in those cases where I applied artiBcial fertilization , see below. 



