294 DIVISION II. — COURSE OF DEVELOPMENT OF FUNGI. 



containing tissue lies originally between each pair of future lamcl ae, which does not 

 however grow with them, bul disappears as the general structure developes and thus 

 completes the delimitation of the surfaces of the lamellae (Fig. 135 g). The dispute 

 is not important and arises perhaps from specific differences. But there is no 

 question that the primordia of the lamellae have no free lateral margin ; their 

 longitudinal margin on the side of the stipe passes into the tissue of the surface 

 of the stipe as continuously as the margin that looks towards the pileus passes into 

 the substance of the pileus. 



The commencement of all the principal parts is followed by a general vigorous 

 growth throughout, and from this time the pileus and the stipe grow more rapidly 

 than the bullous, the contrary having been the case in former stages. The portion of 

 the stipe above the bulbus, hitherto a low narrow disk surrounded by the commencing 

 hymenium, grows into a stout body slightly conically pointed at the apex, and the 

 growth of the young pileus and of the hymenium keeps pace so exactly, that the 

 attachment of the lamellae to the surface of the stipe, which has been already noticed, 

 is not interfered with; the pileus increases in size by superficial extension in the 

 direction of the margin, and the lamellae which arc first formed increase 

 correspondingly in radial length in the same direction, and in breadth in the 

 direction of the stipe. The increase in breadth, however, is comparatively small, so 

 that the marginal portion of the pileus is somewhat curved over and bent downwards 

 (epinastically) in towards the surface of the stipe (Fig. 135 d). Lastly, as the general 

 surface of the hymenium becomes broader towards the margin in consequence of 

 this growth, new lamellae make their appearance between the original ones, being 

 distinguished from them only by the fact that the edge towards the apex of the 

 pileus does not extend as far as the apex, while that towards the margin lengthens as 

 the margin advances ; the later they are formed the greater the distance from the apex 

 of the pileus at which they terminate. All this growth of pileus and lamellae 

 marginwards takes the same course as regards its direction as in the rest of the 

 Agaricineae ; but it does not result in the formation of free margins or even of 

 margins bounded by gaps. The various margins are bounded by undifferentiated 

 hyphal weft with intercalary growth and pass into it without any distinct boundary 

 line, and at the zone of transition the differentiation is constantly spreading over 

 newly formed sections of this weft. 



These processes of growth and differentiation at length reach their limit, and 

 are followed by the final completion of all the parts, the last acropetal elongation of 

 the now tall cylindrical stipe and the hyponastic upward expansion of the pileus 

 —phenomena which again follow in their main features the same course as in the 

 species previously described. There is only one more peculiarity to be observed in 

 the Amaniteae, namely, that the peripheral layer of the stipe into which the edges of 

 the lamellae pass, a thick loosely felted layer, takes no part in the elongation of the 

 inner portion of the stipe which it encloses. It separates everywhere as the 

 elongation proceeds from the surface of the stipe, retaining its connection with it 

 only at the point of insertion of the pileus (Fig. 135 d) ; it also continues at first in 

 connection with the edges of the lamellae, forming a continuous membrane which 

 extends over the whole surface of the hymenium up to the margin of the pileus. As 

 the pileus expands the connection is dissolved, beginning from the margin of the 



