APPENDIX 613 



This species receives its specific name from its peculiar strong odor 

 when handled. 



The archegonia occupy the same position as in Targionia, but the 

 envelope about the sporogonium is much less developed than in the 

 latter. 



The antheridia are formed on very short ventral branches, on the 

 same plants that bear archegonia. 



Lang (6) has made a careful study of this species as well as of a 

 second one which he refers provisionally to C. cavernarum. 



The thallus consists mainly of a single layer of larger air-chambers, 

 bounded below by a single layer of cells, and opening above by well- 

 defined pores like those of Targionia, but there is no trace of the green 

 assimilating filaments found in the latter. In C. fcetidissimum there 

 are several layers of ventral cells in the region of the midrib. The 

 cells of the superficial layer contain a few relatively large chromato- 

 phores, and this is the principal photo-synthetic tissue. 



The archegonia and antheridia closely resemble those of Targionia. 

 As already surmised (see text, p. 70), Leitgeb's suggestion that the 

 antheridium is a single cell has proved incorrect. The early stages 

 of the embryo, as shown by Lang's investigations, resemble the Junger- 

 manniales rather than the Marchantiales. The first two divisions 

 are transverse (as Miss O'Keefe found in Targionia), and the lower- 

 most cells form a sort of haustorium, instead of the massive globular 

 foot found in Targionia. There is a slender but short seta, as in 

 Sphcerocarpus, and except for the presence of a small thickened disc 

 at the summit, the sporogonium more nearly resembles that of Sphcero- 

 carpus than it does Targionia. The wall cells, however, develop 

 thickenings like those found in Targionia, and true elaters are 

 present. 



P. 70. Occasionally receptacles have been found which bear 

 both archegonia and antheridia (see Ernst (i), Cutting (i) ). 



P. 70. Stephani (i) records 200 species of Marchantiaceae, and 

 since his summary was published a number of new species have been 

 described, including several new genera. The Himalayan region is 

 especially rich in these new types (see Kashyap (2) ). 



P. 70. Schiffner, in a recent paper (4), still asserts that Monoclea 

 should be referred to the Jungermanniales ; but the arguments he 

 offers are not very convincing. It may be said, however, in view 

 of the recent work on the Targioniaceae and Pellia (Hutchinson (i) ), 

 that there is a possibility that Monoclea may be in a sense intermediate 

 between the thallose Jungermanniales of the Pellia type, and the 



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