v] 



SPIIAKRIALES 



157 



long hairs around the ostiole, and from the Sphaeriaceae in the habitat and 

 type "f spore. The mycelium is in most cases composed of multinucleate 

 cells, but in Podospora hirsuta the cells are uninucleate (fig. 115), recalling 

 the condition in several species of Chaetomium. 



The commonest type of archicarp is a stout, coiled, septate hypha which 

 soon becomes surrounded by vegetative filaments; it is usually terminal, but 

 is occasionally intercalary, for instance in Sordaria fimicola. Dangeard lias 

 found a straight archicarp (fig. 1 l&) in Sordaria macrospora, and in [868, for 

 S. fimiseda, Woronin described an archicarp with a swollen terminal cell 

 recalling the oogonium of Ilmiuuia granulata. 



Fig. 115. Podospora hirsuta 

 Dang. , archicarp; alter Dan- Fig. 116. Sordaria macrospora Auersw.; a. straight archicarp ; after 

 geard. Dangeard. 



In Sporortnia intermedia the perithecium is initiated by the enlargement 

 of a multinucleate mycelial cell which is often intercalary. It undergoes 

 not only transverse but also longitudinal divisions, forming a pseudoparen- 

 chymatous mass of uninucleate cells (fig. 1 17), with which various neighbouring 

 cells anastomose. The mass thus formed is responsible for the whole contents 

 of the perithecium, though the outer walls may be formed by ordinary 

 vegetative hyphae. In view of this fact it seems doubtful whether the initial 

 cell should here be regarded as an 

 oogonium, that is to say as having at 

 one time had a sexual significance, 

 and not rather as a preliminary stage 

 in tin: development of such a mass <>f 

 hyphae as initiates the apogamous 

 perithecium of Claviceps and its allies. 



In some of the Sordariaceae each 



I11;. 1 1;. Sporormia intermedia Auersw. ; initial 

 spore is surrounded by a layer of cells of perithecium ; after Dangeard. 



