STROPHOMENIA SCANDENS. 109 



a fact that may be responsible for some of the remarkable relations of these glands 

 as described in some other members of the genus and in Rhopalomenia. In the 

 present species these organs are placed side by side beneath the intestine and 

 extend backward from their outlet for a distance equal to at least four times 

 the diameter of the body. Anteriorly they diverge and enter the pharyngeal 

 wall almost directly above the labio-buccal ganglia. Imbedded in muscle and 

 connective-tissue fibres and some of the outlying pharyngeal glands, each canal 

 now bends sharply upon itself (in two individuals) and becoming much more 

 slender courses downward and somewhat backward to enter the pharynx on its 

 ventral side. From the bend to a point close to their outlet these ducts are not 

 only relatively slender but their lumen is of very small size and save in excellently 

 preserved material, as in one of the specimens, is scarcely to be distinguished 

 from the surrounding tissue. They are also composed of remarkably delicate 

 material for in one specimen in which the organs are in a fairly good state of 

 preservation this section of the gland has macerated and disappeared completely, 

 leaving only the short and firmer outlets into the pharynx and the free main 

 part of the gland. 



On the ventral side of the pharynx the glandular portion of each of these 

 organs ceases abruptly and becomes continuous with a canal of somewhat larger 

 calibre composed of small cubical cells, that proceeds almost directly inward 

 and opens ventrally into the pharynx close to the mid line. This terminal sec- 

 tion of the salivary glands is a conspicuous object in sections, but its connection 

 with the adjoining glandular part may be readily overlooked in which case the 

 terminal section appears to be a short diverticulum of the ventral wall of the 

 pharynx while the gland proper seems to unite with the pharynx near the dorsal 

 side. 



In the present species there is in addition to the various divisions of the 

 salivary glands a median diverticulum whose relation to the digestive tract and 

 buccal commissure support the belief that it is a rudimentary radula. The 

 ventral wall of the pharynx forms a shallow pocket and the salivary ducts open 

 into the bottom of this at each side. Exactly in the mid line and immediately 

 above this pouch there is a small posteriorly directed diverticulum of the ventral 

 wall of the pharynx and in two examples the buccal commissure passes dorsal 

 to it (Plate 6, fig. G), the relations being the same as in the radula of P. hawaiiensis 

 for example. Its epithelial lining consists of low cells composed of highly vacu- 

 olated cytoplasm in which the nuclei hold a basal position, but there is no sign 

 whatever of teeth. 



