45 



covering of fine spines has been retained, but it may be easily differ- 

 entiated from all other pupae bearing spines of two sizes in a similar 

 position on account of the large maxillary palpi. 



The family Heliodinidae is included here for the sake of con- 

 venience as it possesses only the cephalic row of spines on the dorsum 

 of the abdominal segments. It is, however, much more nearly related 

 to the Tortricoidea. The families Prodoxidae and Acrolophidae are 

 more nearly related to the Tineidae. Of these the Prodoxidae are 

 undoubtedly the most generalized, retaining more head sutures and a 

 greater number of free segments, in addition to the spines mentioned 

 above. The Acrolophidae are more generalized than the Tineidae in 

 the matter of free segments, but have the appendages firmly soldered 

 to each other and to the body wall. This is probably due to the fact 

 that the larvae are sod-borers and that the pupa works its way to the 

 surface. The families may be separated as follows : 



a. Mesonotum not produced into a long caudal lobe ; mesothorax seldom 

 more than twice the mesal length of the metathorax. 

 b. Abdominal segments 2-7 movable ; dorsum of abdominal segments 

 with a covering of spines on the caudal part ; maxillae more than 



twice as long as the labial palpi Prodoxidae. 



bb. Abdominal segments 3-7 movable ; dorsum of abdominal segments 



never with a covering of spines on the caudal part; maxillae 



shorter than the labial palpi. 



c. Antennae never extending to the caudal margin of the wings; 



wings broadly rounded ; appendages firmly soldered to each 



other and to the body ; a lateral projection never present on 



each side of the tenth abdominal segment Acrolophidae. 



cc. Antennae extending beyond the caudal margin of the wings ; 

 wings pointed; appendages only slightly soldered together 

 and separating at the slightest touch; tenth abdominal seg- 

 ment with a prominent lateral projection on each side end- 

 ing in a spine Tineidae. 



aa. Mesonotum produced into a long caudal lobe ; metathorax never 

 more than one fourth the mesal length of the mesothorax. 



Heliodinidae. 



Family Prodoxidae 



In this family abdominal segments 2-7 are free in both sexes. The 

 head shows the epicranial suture plainly, and dehiscence always takes 

 place on the front of the head along what is apparently the fronto- 

 clypeal suture, at least for a part of the distance, as shown in Figure 

 26. The front bears a prominent chitinized projection armed with 

 two stout teeth. The lateral margin of the eye-piece extends on to 

 the antenna for a very short distance. The appendages are very 



