Superfamily FORMICOIDEA 1439 



nevadensis Cole. Nev. (Kyle Canyon, Charleston Mts.). Ecology: Nests were found in an 



unshaded area in open forests; some were under stones and some were in exposed soil 

 with a scattering of soil around the entrance. 

 Lasius iChthonolasius) nevadensis Cole, 1956. Tenn. Acad. ScL, Jour. 31: 26-27. 5, 9, 6. 

 speculiventris Emery. N. J., Pa. w. to Minn., Iowa, Kans. Ecology: Nests have been found 

 under rocks and in rotting wood either in pastures or wooded areas. 

 Lasius speculiventris Emery, 1893. Zool. Jahrb., Abt. f. System. 7: 637, 641. 5, cJ. 



Taxonomy: Wheeler, 1910. Psyche 17: 237, 242-243. —Wheeler, 1917. Conn. State Geol. and 

 Nat. Hist. Survey Bui. 22: 592. 



Biology: Wilson, 1955. Harvard Univ., Mus. Comp. Zool, Bui. 113: 173. — Kannowski, 1959. 

 Insectes Sociaux 6: 138-141, 153-154 (parasitism on minutus and possible other Lasius 

 spp.). 

 subumbratus Viereck. N. S., Maine w. to Sask., Wash., Oreg., s. to N. Mex., Ariz., Nev. 



Ecology: Nests under stones or rotting logs in meadows and forests. A temporary social 

 parasite. Host: Lasius pallitarsis (Provancher). 

 Lasius umbratus subumbratus Viereck, 1903. Amer. Ent. Soc., Trans. 29: 73. 9. 



Taxonomy: Wheeler, 1910. Psyche 17: 237-239. —Cole, 1942. Amer. Midland Nat. 28: 374. 



Biology: Wheeler, 1917. Amer. Acad. Arts and Sci., Proc. 52: 528. —Wheeler, 1917. Psyche 

 24: 167-176. —Wilson, 1955. Harvard Univ., Mus. Comp. Zool., Bui. 113: 179-180. —Wheeler 

 and Wheeler, 1963. Ants of N. Dak, pp. 191-192. —Gregg, 1963. Ants of Colo., pp. 475-478. 

 umbratus (Nylander). N. S., N. B., Que. s. to Fla. w. to Idaho, Utah, Ariz.; Eiu-asia. Ecology: 



Prefers moist soil and most commonly nests under stones and in or imder rotting logs 



and stumps. May foster subterranean plant lice and mealybugs; occasionally a 



house-infesting ant. A temporary socicd parasite. Host: Lasius alienus (Foerster), L. 



niger (L.), L. neoniger Emery (?). 

 Formica umbrata Nylander, 1846. Acta. Soc. Sci. Fenn. 2: 1048-1050. 9,6. 

 Formica mixta Nylander, 1846. Acta. Soc. Sci. Fenn. 2: 1050-1052. 9. 

 Formica affinis Schenck, 1852. Jahrb. Ver. Nat. Nassau 8: 62-63. 5, 9, S. 

 Formica aphidicola Walsh, 1862. Ent. Soc. Phila., Proc. 1: 310. 5, 6. 

 Lasuis umbratus var. mixto-umbratus Forel, 1874. Nouv. Mem. Soc. Helv. Sci. Nat., p. 48. 



5. 

 Lasius umbratus var. exacutus Ruzsky, 1904. Kasan Univ. Obschchestvo 



estestvoispytatelei Protokoly Zasiedanii, no. 206, p. 15. 5 . 

 Lasius umbratus var. mixto-affinis Ruzsky, 1904. Kasan Univ. Obschchestvo 



estestvoispytatelei Protokoly Zasiedanii, no. 206, p. 15. Nomen nudum. 

 Lasius umbratus var. mixto-bicomis Ruzsky, 1905. Schrift. Naturforsch. (Jesell. Univ. 



Kasan 38: 292. Nomen nudum. 

 Lasius umbratus var. affino-um,bratvs Donisthorpe, 1914. Ent. Rec. 26: 40. 5 . 

 Lasius umbratus var. jrrzewalskii Ruzsky, 1915. Mus. Zool. Acad. Sci. Petrograd 20: 434. 



5. 

 Formicina umbrata distinguenda Emery, 1916. Rend. Accad. Bologna, pp. 64-65. 5,9. 

 Formicina umbrata var. hybrida Emery, 1916. Rend. Accad. Bologna, p. 66. 

 Formicina umbrata var. nuda Bondroit, 1917. Soc. Ent. de France, Bui. 86: 176. 

 Formicina umbrata var. sabularum Bondroit, 1918. Soc. Ent. de France, Bui. 87: 31. 

 Formicina belgarum Bondroit, 1918. Soc. Ent. de France, BuL 87: 31. 5, 9. 

 Lasius bicomis var. citrina Emery, 1922. Soc. Ent. Ital, Bol. 54: 12. 5 . 

 Lasius umbratus var. viehmeyeri Emery, 1922. Soc. Ent. Ital., Bol. 54: 13-15. 5,9. 

 Lasius silvestrii Wheeler, 1928. Lab. Zool. Portici, Bol. 20: 120-121. 9 . 

 Lasius viehmeyeri var. dalmatica Starcke, 1937. Tijdschr. Ent. 80: 53-54. 9. 

 Lasius umbratus var. hirtiscapus Starcke, 1937. Tijdschr. Ent. 80: 43. 9. 

 Lasius umbratus distinguendus var. cereomicans Starcke, 1937. Tijdschr. Ent. 80: 48-49. 



9, 9, 6. 

 Lasius silvestri (!) var. osakana Santschi, 1941. Mitt. Schweiz. Ent. Ges. 18: 278. 9. 

 ChtonolasiusC) affinis var. nyaradi Roszler, 1943. Zool. Anz. 144: 47-48. 5,9. 

 Lasius umbratus epinotalis Buren, 1944. Iowa State Col., Jour. ScL 18: 296, 297. 5 . 



