1812 Hymenoptera in America North of Mexico 



Biology: Cruden, 1972. Evolution 26: 379 (floral relationships). — Cruden, 1972. Madrono 21: 

 511 (floral relationships). 

 nigroclypeata Linsley. Calif. Pollen: Unknown, but visits flowers oi Lasthejiia, Layia 



platyglossa, Marah, Phacelia alba, Plafystemon californkus, Pholistoma racemosa, 

 Raphanus sativa, Tamarix. 

 Andrena nigroclypeata Linsley, 1939. Pan-Pacific Ent. 15: 155. 9, 6. 



Genus ANDRENA Subgenus ONAGRANDRENA Linsley and MacSwain 



Andrena subg. Onagrandrena Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 111. 

 Type-species: Andrena oenotherae Timberlake. Orig. desig. 



The species of this subgenus are chiefly dependent upon the pollen and nectar of several 

 genera within the family Onagraceae, especially Camissonia, Clarkia, and Oenothera. These 

 bees are largely or entirely dark or black in coloration and their behavior is usually closely 

 synchronous with the diurnal flowering regimes of their host plants. Thus some of these bees 

 are matinal species while others are active at the flowers only in the late afternoon and early 

 evening and still others are active at the flowers both in the early morning and again in the late 

 afternoon and early evening hours. 



Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 163-172 (as Melandrena). 



— Linsley and MacSwain, 1956. Pan-Pacific Ent. 32-111-114 (tax. characters, included spp.). 



— Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 117-130 (Oenothera and Clarkia 

 visiting spp.). —Linsley and MacSwain, 1962. Pan-Pacific Ent. 38: 49-52 (new spp. from 

 Calif., Nev., Wyo.). —Linsley and MacSwain, 1963. Pan-Pacific Ent. 39: 189-198 (oenotherae 

 complex). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 5-6 (key to 

 Colorado Desert spp.). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 

 31 (key to Great Basin spp.). —Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 

 33: 64-65 (key to Mojave Desert spp.). — LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 

 284, 287, 290, 292, 315 (tax. characters). —Linsley, MacSwain, Raven and Thorp, 1973. 

 Calif. Univ. Pubs. Ent. 71: 27 (key to cismontane Calif, and Baja California spp.). 



Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 173-185 (nesting habits, 

 flower relationships). —Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 114-121 (nesting 

 habits, floral relationships). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 

 33: 1-58, 6 plates, 6 text figs., 19 tables (comparative behavior of Oenothera visiting spp. 

 on the Colorado Desert and Great Basin). —Linsley, MacSwain and Raven, 1964. Calif. 

 Univ. Pubs. Ent. 33: 59-98, 3 plates, 17 tables (comparative behavior of Oenothera visiting 

 spp. on the Mojave Desert). —MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 

 70: 1-80, 3 plates, 10 text figs., 18 tables (comparative behavior of Clarkia visiting spp. in 

 western U. S.). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 

 1-68, 6 plates, 15 text figs., 10 tables (comparative behavior of Camissonia and Oenothera 

 visiting spp. in cismontane Calif, and Baja California). 

 anograe anograe Cockerell. Calif., Colo. Wyo. Pollen: Collects pollen from Oenothera 



caespitosa, but visits other onagraceous flowers including Aywgra, Oenothera albicaiilis, 

 0. caespitosa var. montana, and may collect pollen from these as well as the large white 

 flowered evening primroses of the subgenus Pachylophis. 

 Andrena anograe Cockerell, 1901. Canad. Ent. 33: 154. 9. 

 Ayidrena micranthophila Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 432. 9, S. 



Taxonomy: Cockerell, 1934. Amer. Mus. Novitates 697: 2 (tax. characters). —Linsley and 

 MacSwain, 1955. Pan-Pacific Ent. 31: 166 (synonymy, geogr. records). —Linsley and 

 MacSwain, 1961. Pan-Pacific Ent. 37: 126 (changed status). —Linsley, MacSwain, and 

 Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (tax. characters). 



Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 39 (floral 

 relationships). 

 anograe knowltoni Linsley and MacSwain. Utah. Pollen: Unknown, but visits flowers of 

 Camissonia scapoidea scapoidea, Staiileya pinnata. 

 Andrena (Onagrandrena) anograe knowltoni Linsley and MacSwain, 1961. Pan-Pacific 

 Ent. 37: 126. 9 . 



