Superfamily APOIDEA 2009 



Type-species: Apis florisomnis Linnaeus. Monotypic. {=Apis maxillosa Linnaeus). 

 Gyrodroma Thomson, 1872. Hym. Scand., v. 2, p. 259. Preocc. 



Type-species: Heriades nigricoimis Nylander. Desig. by Cockerell, 1925. 



Nearctic species of this genus visit flowers of Hydrophyilaceae, especially the genera 

 Phacelia and Eriodictyon, and possibly collect pollen exclusively from these flowers. 



Revision: Michener, 1938. Pan-Pacific Ent. 14: 36-45. 

 bernardinum Michener. South. Calif. Pollen: Unknown, but visits flowers oi Nemophila 

 integrifolia, N. vienziesii, N. spatulata, Phacelia davidsonii. 

 Chelostoma bernardinum Michener, 1938. Pan-Pacific Ent. 14: 40. 9, 6. 

 californicum Cresson. Calif.; Mexico (Baja California). Pollen: Apparently oligolectic on 

 flowers of Phacelia including P. ciliata, P. davidsonii, P. distans, P. hispida, P. 

 imbricata, P. platyloba, P. ramosissima, but visits other flowers presumably for nectar 

 including Cryptantha inter)nedia, C.flaccida, Eriodictyon californicum, E. trichocaiyx, 

 Hesperochiro)i califoriiicuni, Layia platyglossa, Nemophila vienziesii, N. integrifolia, 

 Plagiobothrys )iothofi(lvHs, Salvia carnosa. 

 Chelostoma californicum Cresson, 1878. Amer. Ent. Soc, Trans. 7: 108. 6. 

 Heriades albicinctunt Provancher, 1895. Nat. Canad. 22: 190. 9,6. 

 Heriades odontura Cockerel!, 1902. South. Calif. Acad. Sci., Bui. 1: 139. 6. 

 Roberfsonella dolichosonia Cockerell, 1922. Ann. and Mag. Nat. Hist. (9) 10: 456. 9. 

 cockerelli Michener. South. Calif. Pollen: Apparently oligolectic on flowers of Enodictyon 

 including E. califoniicnm, E. crassifoliutn, E. trichocaiyx, but visits other flowers 

 including Clarkia, Cryptantha intermedia, Gilia exilis, Lotus scoparius, Mimulus 

 fremontii, Nania parryi, Navarretia heterodoxa, Penstemon, Phacelia distans, P. 

 tana ceti folia, Tricbostema lanatnm, T. pariskii. Verbena lasiostachys, Viguiera 

 deltoides. 

 Chelostoma niinutuni cockerelli Michener, 1938. Pan-Pacific Ent. 14: 43. i, 9. 

 incisulum Michener. Calif. Pollen: Apparently oligolectic on flowers of Phacelia including P. 

 distans, P. platyloba, but also visits flowers oi Ceanothus, Cryptantha, Gilia tricolor, 

 Hesperochiron californicus, Lasthenia chrysostoma, Linanthus aureus. 

 Chelostoma >ninutu))i incisulum Michener, 1938. Pan-Pacific Ent. 14: 44. 6. 

 marginatum incisuloides Michener. Cent. Calif. Pollen: Apparently oligolectic on flowers of 

 Phacelia including P. platyloba, but also visits other flowers including Amsinckia, 

 Cryptantha, Eriodictyon californicum, Nemophila pulchella, Penstemon laetus. 

 Chelostoma marginatu))i incisuloides Michener, 1954. Kans. Ent. Soc, Jour. 27: 69. 6, 9. 

 marginatum marginatum Michener. South Calif. Pollen: Apparently oligolectic on flowers of 

 Phacelia including Phacelia davidsonii, P. distans, P. hispida, P. minor, but also visits 

 other flowers including Allium pannim, Cryptantha intermedia, Eriodictyon 

 crassifolium, Lomatium dasycarpum, Nemophila pulchella, Rharnnus crocea, Rhus 

 trilobata. Salvia columbariae, S. mellifera. 

 Chelostoma miyiutum marginatum Michener, 1938. Pan-Pacific Ent. 14: 44. 6, 9. 

 minutum Crawford. Wash., Oreg., Utah, Nev., Calif. (Sierra Nevada and San Bernardino Mts.), 

 Idaho. Pollen: Presumably oligolectic on Phacelia including P. californica, P. davidsonii, 

 P. distans, P. heterophylla, P. imbricata, P. liyiearis, P. ramosissi^na, but also visits 

 other flowers including Allium parvum, Calochortus, Cryptantha, Draperia systyla, 

 Erigeron divergens, Eriodictyon crassifolium, E. trichocaiyx, Eriophyllum lanatum, 

 Erysimum asperum, Heuchera micrantha, Lepidium virginicum, Lotus argophyllus, 

 Mimulus fremontii, Nemophila menziesii, Penstemon spectabilis, Polemonium, 

 Potentilla glandulosa, Sidalcea maivaeflora. 

 Chelostoma minuta Crawford, 1916. Insecutor Inscitiae Menstruus 4: 102. 9. 

 phaceliae Michener. Wash., Oreg., Calif., Nev. Pollen: Apparently oligolectic on Phacelia 



including P. californica, P. distans, P. heterophylla, P. hispida P. imbricata, P. linearis, 

 P. ramosissima, P. tanacetifolia, but also visits other flowers including Allium 

 dichlamydium, Brodiaea laxa, Calochortus, Clarkia dudleyana, C. pulchella, C. 

 rhomboidea, Cryptantha intermedia, Erigeron foliosus var. stenophyllus, Eriodictyon 

 californicum, E. crassifolium, Gilia exilis, Horkelia bolanderi, Lepechinia calcycina. 



