2154 Hymenoptera in America North of Mexico 



wheeleri Cockerell. Mich, and N. Dak., south to La., Tex. and Ariz. Pollen: Oligolege of 

 Compositae, especially Gaillardia, Helianthns and Rudbeckia, visits flowers of 

 Anthemis cotula, Asclepias tuberosa, Cleome serrulata. Coreopsis, Echinacea, 

 Engelmannia bipinnatifida, Gaillardia pulchella, Grindelia squarrosa, Helianthus 

 annuus, H. debilis, H. petiolaris, Opuntia, Petalostemon purpiireum, Ratibida 

 columnaris, Rudbeckia bicolor, R. grandiflora, R. hirta, Silphium asperimum, 

 Sphaeralcea. 

 Melissodes wheeleri Cockerell, 1906. Amer. Ent. Soc, Trans. 32: HI. 9. 



Taxonomy: Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 367. 6. 



Genus MELISSODES Subgenus PSILOMELISSODES LaBerge 



Melissodes subg. Psilomelissodes LaBerge, 1956. Kans. Univ. Sci. Bui. 37: 1173. 

 Type-species: Melissodes intorta Cresson. Monotypic and orig. desig. 

 intorta Cresson. Kans., Tex. Pollen: Probably dependent upon the pollen of the Malvaceous 

 genus Callirhoe including C. digitata, C. involucrata, C. leiocarpa, but also visits 

 flowers of Asclepias latifolia, Gaillardia, Rorippa sinuata. 

 Melissodes intorta Cresson, 1872. Amer. Ent. Soc, Trans. 4: 278. 6. 

 Melissodes wickhami Cockerell, 1906. Amer. Ent. Soc, Trans. 32: 112. 9. 



Taxonomy: LaBerge, 1963. Nebr. Univ. State Mus., BuL 4: 234 (geogr. and floral records). 

 Genus MELISSODES Subgenus CALLIMELISSODES LaBerge 



Melissodes subg. Callimelissodes LaBerge, 1961. Kans. Univ. Sci. Bui. 42: 294. 

 Type-species: Melissodes iupina Cresson. Orig. desig. 

 ablusa Cockerell. Calif. (Millbrae, Bolinas). 



Melissodes metenua ablusa Cockerell, 1926. Pan-Pacific Ent. 3: 85. 9. 

 clarkiae LaBerge. Oreg., Calif. Pollen: Probably oligolectic on Clarkia including C. amoena, C. 

 biloba, C. cylindrica, C. dudleyana, C. gracilis albicaulis, C. speciosa, C. uyigtiiculata, C. 

 William sonii. 

 Melissodes (Callimelissodes) clarkiae LaBerge, 1961. Kans. Univ. Sci. Bui. 42: 315, figs. 

 50-53. 9, cJ. 



Biology: MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 29-31 (floral 



relationships). 

 coloradensis Cresson. N. C, Ohio, Ind., Ill, Wis., Mo., Nebr., Kans., Wyo., Colo., N. Mex., Ariz., 



Utah, north. Calif. Pollen: Oligolege of Compositae, with Helianthus including H. 



annuus, H. atrorubens, H. divaricatus, H. grosse-serratus, H. mollis, H. petiolaris, H. 



scaberrimus and H. tuberosus as the primary source of pollen, but also visits flowers of 



Cirsium discolor, C. lanceolatum. Coreopsis tripteris, Heliopsis helianthoides, H. laevis, 



Liatris pycnostachya, Lythrum alatum, Rudbeckia laciniata, Silphium integrifolium, S. 



laciyiiatum, S. perfoliatum, S. terebinthinaceum, Solanum, Solidago serotina. Verbena 



hastata, V. stricta, Vemonia baldwini interior, V.fascictilatum. 

 Melissodes coloradensis Cresson, 1878. Acad. Nat. Sci. Phila., Proc 30: 200. 9, c?. 

 composita Tucker. Mont, south to Ariz., N. Mex. Ecology: Nests in ground individually, but 



many individuals use a communal entrance. Parasite: Triepeolus helianthi Robt.? Pollen: 



Unknown, but visits flowers of Grindelia squarrosa, Haplopappus gracilis, Thurberia 



thespesioides. 

 Melissodes Iupina composita Tucker, 1909. Kans. Acad. Sci., Trans. 22: 281. 3. 



Biology: Hurd and Linsley, 1959. Ent. News 70: 141-146 (nest). 

 glenwoodensis Cockerell. N. Dak. south to N. Mex., west to Wash, and southern Calif. Pollen: 

 Oligolege of Compositae, especially Chrysothamnus including C. nauseosus coiisiinilis, 

 C. n. mojavensis, C. n. speciosus, C. viscidiflorus typicus, but visits other flowers 

 including Cirsium, Grindelia squarrosa, Haplopappus vemonioides, Helianthus 

 petiolaris, Solidago trinervata. 

 Melissodes glenwoodensis Cockerell, 1905. Ann. and Mag. Nat. Hist. (7) 15: 522. 9. 



Taxonomy: Cockerell, 1906. Amer. Ent. Soc, Trans. 32: 107. 



