378 



THE BASIDIOMYCETES 



expands into a pileate structure, so that the glebal surface is more 

 extensive than in Mutinus. The culmination of development is 

 reached in Dictyophora, which is similar to Phallus but has in 

 addition a net-like veil or indusium, originating between the 

 pileus and the stalk and encircling the stalk [Atkinson (1911)]. 

 There is great variation among the different species in the de- 





Fig. 147. Various Phallales. A. Clathnis cancellams. B. Si?nblum sphaero- 

 cephalum. C. A?itbiirus borealis. D. Mut'nms caninus. E. Ithyphallus 



impudiciis. 



gree of development of the veil and the nature of the pileate 

 portion of the receptaculum. Tt has been found [Burt (1897)] 

 that the rapid elongation of the receptaculum, which may shoot 

 up to its mature length overnight, results from quick growth 

 (cellular enlargement) and absorption of water, and the process 

 is accompanied by a depletion of glycogen. 



The Clathraceae represent another developmental series, in 

 which the gleba occupies an internal position in relation to the 

 receptaculum. In Clathrus the receptaculum is an ovoid, hollow, 

 lattice-like structure, seated on the fragments of the ruptured pe- 

 ridium. The gleba occupies the entire interior surface of the 

 interlacing arms. In Simblum [Conard (1913)] the basal portion 

 of the receptaculum constitutes a long stalk, w^hose apex is clath- 

 rate, so that the whole fruiting body resembles a miniature 

 Clathrus on a stem. 



