134 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL.5 



the general zoogeographic position of the American west coast (cf. 

 Ekman, 1935). Lysiosquilla digueti Cout. from the Gulf of California 

 has as its nearest relative L. vicina Nobili from the Red Sea and the 

 Philippines. Pseudosquilla lessonii Guerin (Chile to California) is re- 

 lated to P. dofleini Balss from Japan and Hemisquilla stylifera (H.M.- 

 Edw.) (Chile to California) has otherwise been found only in New 

 South Wales." 



The stomatopods described as new or which have been given new 

 names in this paper fit comfortably into such a distribution pattern. 

 There can be no question regarding the Atlantic affinities of the several 

 Gonodactylus forms, G. bahiahondensis, G. festae lalibertadensis, and 

 G. stanschi; or of Squilla bigelowi, S. hancocki, and S. tiburonensis. The 

 Indo-Pacific ties of Lysiosquilla maccullochae to L. latifrons (Japan, Aus- 

 tralia, New Zealand), of Pseudosquilla veleronis to P. pilaensis (China, 

 Indian Ocean, Red Sea), and of Squilla swetti to 5. fasciata (Japan, 

 China, and Indian Ocean) or S. miles (Australia), are very much less 

 significant, and therefore throw into all the more striking relief the 

 near identity of Squilla hildebrandi from Panama to S. hieroglyphica 

 which, although without definite locality, is most assuredly of Indo- 

 Pacific origin. 1® 



Whether Squilla hildebrandi is a relict, or has evolved from a relict 

 of a formerly widespread Pacific fauna, or is the descendant of a 

 member of a drift fauna transported to the shores of America by some 

 current similar to or stronger than the relatively weak Pacific equatorial 

 countercurrent, is impossible to say. Of more than passing interest in this 

 connection was the discovery of the Indo-Pacific coral gall crab, Hapa- 

 locarcinus marsupialis, at Port Utria, Colombia, and in the Secas Islands, 

 Panama, by the Hancock Expedition of 1935, ^^ and that of the first 

 Thalamita from the Western Hemisphere, T. roosevelti, at Clipperton 

 Island during the Presidential Cruise of 1938.20 At the same time, at 

 Clipperton Island, I also found a terrestrial amphipod almost specifically 

 identical with one described from the Marquesas Islands. A number of 

 Polynesian mollusks, or near relatives of such shells, have been reported 



18 S. hieroglyphica in turn is closely related to S. laevis Hess from Australia 

 (Kemp, Mem. Indian Mus., Vol. 4, No. 1, p. 49, pi. 3, figs. 35-37, 1913). The other 

 Indo-Pacific species mentioned are also to be found in Kemp. 



19 Exploration and Field-work of the Smithsonian Inst, in 1935, p. 34, fig. 36, 

 1936, and Rathbun, Bull. U.S. Nat. Mus., No. 166, p. 259, pi. 79, figs. 3-9, 1937. 



20 Smithsonian Misc. Coll., Vol. 98, No. 6, p. 16, fig. 2, 1939. 



