94 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 12 



smooth to somewhat crispate, obtuse, to about 3-4 cm broad, 6-10 cm 

 long, the margin entire to slightly sinuate with occasional broad, some- 

 what pointed, lobes; reproductive cells usually solitary, sometimes 3-4 

 together, spherical or mutually compressed in surface view, 57-70 fx. diam. 



Especially in no. 34-155 the blades show traces of a slight midrib for 

 1-3 cm, above which it disappears; Spatoglossiim is customarily defined as 

 lacking a midrib, but Howe (1914, p. 68) describes one in S. crispata. 

 The reproductive cells observed look like immature tetrasporangia, which 

 in this family commonly divide just before dispersal. Scattered oogonia 

 are described for the genus. Some individuals are much more minutely 

 divided than others, with segments 1-2 cm broad, while in one instance 

 an ultimate segment (except for two small proliferations) was 6 cm 

 broad, 23 cm long, and no. 34-386 showed one undivided fragment over 

 9 cm broad, 50 cm long, lacking both base and tip of the segment, indicat- 

 ing that this species reaches noble proportions in deep water. This was 

 presumably a tetrasporangial specimen, many sporangia having germinated 

 in situ, so that small folioles were scattered abundantly over the surface. 

 The sporangia became 60-90 /x diam. before dividing; they seldom if ever 

 became normal tetrads, but produced small leaflets directly. These quickly 

 developed a row of growth initials at the forward margin. The plants 

 were commonly perforate, and near the base almost netlike, but this may 

 not be a natural state. Structurally, the blade developed four layers of 

 cells in the medulla, but the sections did not swell well enough to afford 

 a measurement of thickness. The cortex, of one layer, did, however, expand 

 perfectly; its cells were square to brick-shaped, and about 30-45 /i in 

 surface view, 30 p. in thickness. 



Ecuador: Archipielago de Colon, dredged at sta. 155 off Tagus 

 Cove, I. Isabela, no. 34-181 (?), 15 Jan. 1934. Ibid., infrequent as 

 dredged from 27 meters off Post Office Bay, I. Santa Maria, no. 34-294 

 (TYPE), 19 Jan. 1934. Ibid., occasional at 55 meters' depth, no. 34- 

 386B, 29 Jan. 1934. Ibid., fragments dredged from 37 meters' depth 

 off a rocky bottom, I. Espanola, Schmitt no. 362D-35, 19 Dec. 1934. 



DIGTYOPTERIS Lamouroux, 1809 



Plants small to large and coarse, sometimes entangled, generally erect 

 and bushy, the segments flat, strap shaped to membranous, costate, dichot- 

 omously to alternately branched ; growing from a small group of apical 

 initial cells ; forming the reproductive cells in linear or rounded sori. 



