334 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 21 



Color in life: Carapace and legs uniformly deep olive buff. A large 

 patch of bright madder brown on anterior branchial and hepatic regions. 

 Eyes brilliant flame scarlet. Chelae and propodi of ambulatory legs lightly 

 shaded with red orange, proximal portion of ambulatory legs light Rus- 

 sian blue. Abdomen and sternum pale mallow pink. Pterygostomian 

 region and maxillipeds madder brown. (Petersen) 



Habitat: Broken shell, broken coral, Albatross. (Rathbun) Sand 

 bottom, Zaca. (Crane) From coral, Askoy. (Garth) The 28 Velero 

 stations for which data on bottom type are available give a primary 

 breakdown as follows: rock 6, sand 9, mud 3, organic (coral, nullipore, 

 and coralline) 10. A secondary breakdown shows shell present with the 

 sand in 2 cases, shell or nullipore present with the rock in 3 cases, and 

 shell or sand present with the mud in all cases. The only bottom on 

 which Lissa tuberosa does not occur would appear to be unadulterated 

 mud. Two specimens from Panama carry a rhizocephalan parasite. 

 Living Bryozoa are frequently recorded as encrusting organisms. 



Depth: Shore to 47 fathoms. 



Size and sex: The series includes males from 5.3 to 12.3 mm, females 

 from 5.8 to 14.8 mm, ovigerous females from 5.8 to 14.5 mm. As in the 

 case of Lissa aurivilliusi, specimens from the Gulf of California are 

 larger than average, the 14.8 mm female being from the Gulf, whereas 

 the largest from the Bay of Panama is 13.5 mm. None are as large as 

 the 16.9 x 15.8 mm male holotype. 



Breeding: Ovigerous females were encountered off Lower California 

 in March, in the Gulf of California in February and March, off the 

 Mexican mainland in March, in the Revilla Gigedo Islands in June, 

 off Panama in February, and off Colombia in January. The 5.8 mm 

 specimen from Cocos Island, Costa Rica, is unusually small to be gravid 

 and carries a correspondingly small number of large-sized eggs. The 

 smallest ovigerous female from a mainland station measures 9.3 mm. 



Remarks: With the exception of the single specimen taken by the 

 Askoy at Utria Bay, Colombia (Garth, 1948), Lissa tuberosa has been 

 unknown outside of the Gulf of California. As a result of the work of 

 the Velero HI and Velero IV, the range of the species has been ex- 

 tended northward to include Magdalena Bay, Lower California, and 

 Tortuga Island, Gulf of California; westward to include Clarion Island 

 of the Revilla Gigedo Islands; and southward to Octavia Bay, Colom- 

 bia. It is not known to occur in the Galapagos archipelago. 



