354 AQUATIC PHYCOMYCETES 



Since the apophysis of the sporangium is epibiotic the species cannot 

 be placed in Phlyctochytrium as here defined. The resting spores Dan- 

 geard first described for this species (1886a) differ from those he de- 

 scribed in the 1889 paper in being rough-walled and not apophysate, 

 rather than spiny-walled and apophysate. There is a strong possibility 

 that two different organisms are involved. 



Dangeard pointed out that the position of the apophysis (whether 

 sessile or borne on a stalk) depended upon the proximity of the germi- 

 nating zoospore to the host cell. Thus, if the zoospore was at some 

 distance from the Euglena a stalk was formed, otherwise the apophysis 

 was sessile. He distinguished two forms of his fungus: (1) "Chytridium.' 

 Form" in which the apophysis was endobiotic, and (2) " Rhizidium- 

 Form" in which it was epibiotic. It is possible the "Chytridium-Form" 

 is in reality another fungus. As understood here, the species forms 

 typically an epibiotic apophysis. 



Minden identifies with Dangeard's organism a fungus discussed 

 by Schenk (1858a: 246) as Chytridium euglenae Braun, and uses the 

 binomial Phlyctochytrium euglenae (Schenk) Schroeter. Schenk, how- 

 ever, was unquestionably dealing with Polyphagus euglenae. Further- 

 more, Schenk's fungus, though described in the course of a discussion 

 of Braun's genus Rhizidium, was always referred to by him as Chytridium 

 euglenae and not Rhizidium euglenae, as Minden implies. It is interesting 

 to note that Minden (1915:383) in describing Polyphagus euglenae 

 asserts that Schenk's fungus belongs in Polyphagus. See also Chytridium 

 euglenae Braun, page 532. 



Dangeard's fungus is considered by Minden to be synonymous with 

 Saccomyces dangeardii (S. endogenus). 



Braun's Chytridium euglenae, collected by von Siebold and Meissner, 

 is probably identical with the present species but is too little known to 

 be considered here. 



BLYTTIOMYCES Bartsch, emend. 

 Mycologia, 31: 559. 1939 



Thallus epi- and endobiotic, monocentric, eucarpic, consisting of the 

 epibiotic rudiment of the sporangium, provided with an apiculus, which 



