HEMIASCOM YCETES 



143 



are formed directly from the sprout cells, without this detour (Mangenot, 

 1922). Both species ferment sucrose and a series of monosaccharides. 



In two other species which terminate the isogamous development, 

 Endomyces javanensis and E. capsularis (Saccharomycopsis capsularis), 

 the copulation branches have entirely disappeared (Fig. 88, 3 to 5). 

 According to cultural conditions, either the hyphal or sprouting condition 

 preponderates. At the end of the hyphae, there arise by swelling of cells 

 or by lateral sprouting, at times also intercalary, four-spored asci whose 

 spores are divided into two unequal halves by annular thickenings 

 (Guillermond, 1909). Both species have been isolated from earth, 

 the former in Java, the latter in the Swiss Alps. In both, the fermenta- 

 tive ability is small. 



Fig. 88. — Endomyces Lindneri. 1, 2. E. capsularis 3 to 5. Development of asci. (1, 

 2 X 470; 3 to 5 X 500; after Mangenot, 1922, and Guillermond, 1909.) 



The second, heterogamous series is connected to the Eremascus fertilis 

 type through Endomyces Magnusii, found in the slime-flux of trees. Its 

 hyphae are generally multinucleate (2 to 8); in the growing hyphal tips 

 the number of nuclei may mount to 50 (Fig. 89, 1), in weak hyphae it 

 may sink to one. In contrast to Eremascus fertilis, and like Endomyces 

 fibuliger, the hyphae divide easily into oidia; naturally these are generally 

 multinucleate, rarely uninucleate, with a tendency toward the uninu- 

 cleate condition. Often they thicken the wall and become gemmae 

 (Fig. 89, 14); hence under certain conditions, e.g., in Raulin's solution, 

 the whole culture may disintegrate after two weeks into oidia which 

 change into gemmae. 



Under favorable conditions the oidia, as in E. fibuliger, may develop 

 to sprout mycelia. This, however, does not take place by the inde- 

 pendent development of the small outgrowths of the mother cell to sprout 



