Guilliermond - Atkinson 



— 200 — 



Cytoplasm 



pregnate it with osmium. After this treatment, the cells show at 

 one of the poles corresponding to the direction of centrifugal force, 

 an accumulation of chondriosomes and starch-bearing plastids. At 

 the opposite pole are accumulated in order, the lipides, the vacuolar 

 sap and the osmiophilic platelets which consequently seem to be 

 lighter. But the figures given by the authors are not very con- 

 vincing and it seems that the substance which was affected by 

 the centrifugal force corresponds only to starch-containing plas- 

 tids and that the so-called osmiophilic platelets represent vesicu- 

 lated chondriosomes. 



It has been seen that with osmic methods KlYOHARA obtained 



analogous figures (vesicles) which he 

 interprets as normal forms of the plas- 

 tids. This author, starting with a false 

 premise in the form of a defective ob- 

 servation of living material in which 

 he saw only vesiculated chondriosomes, 

 concluded that mitochondrial methods 

 alter the chondriosomes, whereas the 

 Golgi methods preserve them in their 

 vesicular form (C/. p. 91). Our 

 work, however, has furnished proof 

 that these so-called osmiophilic plate- 

 lets are none other than vesiculated 

 chondriosomes and plastids. Finally, 

 Weier, not being able to find in plants 

 a Golgi apparatus independent of 

 formations already known, and having 

 succeeded, in Polytrichum commune, 

 in impregnating large plastids by Golgi 

 methods, came to the conclusion that 

 the Golgi apparatus is represented in 



paratus (G) and chondriosomes plant CCllS by thc plastldS (C/. PP. 91, 

 (M). These Golgi elements are in t,^^ rm . . • • i • i 



reality plastids. 92). This opmion IS howcvcr, mad- 



missible, for the reason that the plas- 

 tids are a variety of chondriosomal elements, belonging to chloro- 

 phyll-containing cells, and in direct relation with photosynthesis 

 which characterizes these cells, and they are not found in fungi. 

 Besides, the ordinary chondriosomes are impregnated by Golgi 

 methods just as well as the plastids. 



It must be added that Gicklhorn, studying large spherical 

 bodies which are found localized in the vacuoles of epidermal cells 

 of Iris, noticed that under the influence of osmic acid they blacken 

 and become a spongy structure, then are transformed into a net- 

 work which looks like Golgi material. This worker thinks, there- 

 fore, that it is to structures of this nature that the Golgi formations 

 must be attributed. The work of one of our students, Reilhes 

 seems to have demonstrated that these bodies are composed of 

 a phytosterol. Liebaldt has recently supported this same opinion. 

 It is possible that analogous formations have been described in 



Fig. 137. — Allium Cepa. Drew's 

 figures of the Golgi apparatus in 

 the root. 1, 2, meristem; only 

 chondriosomes dififerentiated. 3, 4, 

 differentiating cells with Golgi ap- 



