THE MALE GAMETOPHYTE 



171 



D E F 



Fig. 104. Development of male gametophyte of Scirpvs paluster. A, telophase 

 of Meiosis II resulting in formation of four microspore nuclei. B, three of micro- 

 spore nuclei pushed to one end of pollen grain; functioning nucleus in center. C, D, 

 functioning nucleus dividing; remaining three nuclei in process of degeneration. 

 E, pollen grain, showing vegetative and generative cells. F, generative cell dividing 

 to form two male cells. (After Piech, 192S.) 



Embryo-sac-like Pollen Grains. In 1898 Nemec noted that in 

 the petaloid anthers of Hyacinthus orientalis the pollen grains some- 

 times form large eight-nucleate structures showing a surprising 

 resemblance to embryo sacs. He believed that they arose as the 



