THE MALE GAMETOPHYTE 175 



Shortly after the publication of Stow's papers, Naithani (1937) 

 found embryo-sac-like pollen grains in the variety "Yellow Hammer" 

 whose bulbs had been treated for early flowering. He confirms 

 Stow's observations regarding the mode of development of these 

 abnormal pollen grains, but believes their formation to be a tempera- 

 ture effect and not the result of a liberation of necrohormones. 

 According to him, the degeneration of the other pollen grains is not 

 the cause but the effect of a hypertrophied growth of the more 

 favored ones, which use up all the available food for their own 

 growth. 



More recently, pollen -embryo sacs with 8 and 16 nuclei have also 

 been observed in another plant, Orniihogalum nutans (Geitler, 

 1941). Those with 8 nuclei showed the typical embryo-sac-like 

 organization, but, contrary to Stow, Geitler interprets the three 

 cells at the exine end of the pollen grain as the equivalents of anti- 

 podals and the other three as equivalents of the egg and synergids. 



References 



Anderson, L. E. 1939. Cytoplasmic inclusions in the male gametes of Lilium' 



Amer. Jour. Bot. 26: 761-766. 

 Banerji, I., and Gangulee, H. C. 1937. Spermatogenesis in Eichhornia crassipes 



Solms. Jour. Indian Bot. Soc. 16: 289-296. 

 and Haldar, S. 1942. A contribution to the morphology and cytology 



of Monochoria hastaefolia Presl. Proc. Indian Acad. Sci. Sect. B. 16: 91-106. 

 Barber, H. N. 1942. The pollen grain division in the Orchidaceae. Jour. Genet. 



43: 97-103. 

 Beatty, A. V. 1943. The division of the generative nucleus in Eschscholtzia. 



Amer. Jour. Bot. 30: 378-382. 

 Benetskaia, G. K. 1939. Observations in vivo des gametes males dans les tubes 



polliniques d'Asclepias cornutii Decsn., de Vinca major L. et de Vinca minor L. 



Jour. Bot. de l'U.R.S.S. 24: 273-281. 

 Billings, F. H. 1934. Male gametophyte of Atriplex hymenelytra. Bot. Gaz. 



95: 477-484. 

 Brumfield, R. T. 1941. Asymmetrical spindles in the first microspore division 



of certain angiosperms. Amer. Jour. Bot. 28: 713-722. 

 Capoor, S. P. 1937a. Contributions to the morphology of some Indian Liliaceae. 



II. The gametophytes of Urginea indica Kunth. Beihefte bot. Centbl. 56 A: 



156-170. 

 . 19376. The life history of Holoptelea integrifolia Planch. Beihefte bot. 



Centbl. 57A: 233-249. 

 Cooper, D. C. 1935. Microsporogenesis and the development of the male 



gametes in Portulaca oleracea. Amer. Jour. Bot. 22: 453-459. 

 . 1936. Development of the male gametes in Lilium. Bot. Gaz. 98: 



389-177. 



