Asplenium ASPLENIACEAE Athyrium 



tall; fr. V-XI; leaves bipinnate to tripinnate. Eastern Asia; locally in An., Che., 

 Ki., Ku. Crevices of rocks. 



2. Athyrium Roth ffiWfcM ( T 'i Pa0 Chiieh Shu; Hoof Indusiate Fern Genus) 

 Medium to tall terrestrial ferns; leaves often variable in size and degree of cutting, 

 pinnate to decompound: many species bearing spines on upper surface along the sulcate 

 costae: stipes green or stramineous: scales on stipe, rachis and rachillae usually few 

 and thin, not clathrate: sori linear, more or less curved and crossing the veinlets, 

 sometimes reniform; indusia sometimes fugacious: veins free, usually ending in clavate 

 tips. About 180 species. (From the Greek, meaning "without shield," the indusia 

 being on only one side of the veinlets.) 



Key to the Species 



a. Indusia mostly dryopteroid, i.e. reniform. 

 Leaves pinnate to bipinnate. 



Pinnae lanceolate or falcate; apices attenuate 1. A. sheareri 



Pinnae oblong ; apices rounded 2. A. macrocarpum 



Leaves tripinnatifid ; pinnae 15-30 cm. long 3. A. viridifrons 



aa. Indusia athyroid to diplazoid, i.e. slightly curved and crossing 

 the veinlet to completely doubled, 

 b. Lamina gradually attenuate towards apex. 



c. Lamina pinnate; pinnae falcate 4. A. wardii 



cc. Lamina more than once divided; basal pinnules appressed 

 against rachis; with spines along costa on upper sur- 

 face. 



d. Leaves bipinnatifid; lower pinnae shortened 5. A. yokoscense 



dd. Leaves tripinnatifid to tripinnate. 



Lower pinnae 20-30 cm. long 8. A. giganteum 



Lower pinnae 4-12 cm. long. 



Lower pinnae alternate; stipe shorter than lamina 6. A. goeringianum 

 Lower pinnae opposite; stipe longer than lamina 7. A. devolii 

 bb. Lateral pinnae much longer than the pinnate or bipinnatifid apex, 

 making an abrupt break about 1/4 or less of the way from 

 the apex of the leaf. 



Leaves tripinnatifid to tripinnate 9. A. niponicum 



Leaves bipinnate. 



Pinnules acute 9. A. niponicum 



Pinnules obtuse 4. A. wardii 



1. Athyrium sheareri (Bak.) Ching (Ching 3:108). 30-50 cm. tall; fr. IV-X; sori 

 small; indusia caducous. China, Japan; locally in Ki. Stream banks. 



2. Athyrium macrocarpum (Bl.) Bedd. (Bd. F. 165). 25-50 cm. tall; fr. VII-IX; 

 indusia large; margins incised. China, Japan; locally in An., Ki. Damp rocky 

 caves. 



3. Athyrium viridifrons Mak. (A. okuboanum Mak.) (Mak. F. 936; DeV. F. 97). 

 80-120 cm. tall; fr. VI-X; sori small, round. China, Japan; locally in An., Ki. 

 Stream margins. 



4. Athyrium wardii (Hook.) Mak. {Asplenium wardii Hook.) (H. S. F. 217). 40-60 

 cm. tall; fr. V-X; indusia not crossing veinlets. China, Korea, Japan; locally in 

 An., Ki. Wooded hillsides. 



5. Athyrium yokoscense (Fr. & Sav.) Christ (Mak. F. 937). 20-80 cm. tall; fr. VII- 

 X; lower pinnae deflexed. China, Japan; locally in An., Che., Ki. Hillsides. 



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