395 



T. T. Bajinister and M. J. VroomeLn 



increment of short-wave illumination will be divided equally 

 between both reactions. Hence, at this balance point, and for 

 all higher short-wave illuminations, E and D have fixed maLximum 

 values. 



From equations 3 through 6, expressions for E and D are easily 

 obtained, either in terms of i and i , or p^ and p^. Thus, for 

 i, less than balancing 

 1 



E = l+(b-2)i^/ai2 = l+(2b-l)p^/p2 (T) 



D = (b-2)i^ = (2b-l)p3^ (8) 



and, for just balancing and higher short-wave illuminations 



E = l/2a (9) 



D = (l/2a - l)p (10) 



2 



Also, from equation 6, is obtained an expression for the ratio 



p,/pp at the balance point 



2a-l 



p^/p^ = 2a(l-2b} (11) 



Separately packaged pigment model . Here, no energy transfer 

 is assumed possible between the two systems. As a result, 

 absorbed short-wave quanta, like far- red quanta, are presumed 

 to be divided in a fixed ratio between the two photoreactions, 

 the preponderance of short-wave quanta acting in the short-wave 

 system. Then the rate of photosynthesis in short-wave light is 

 limited to the rate of the far-red reaction, giving in place of 

 equation k, 



^ - (i-».)i. 



P, = 



(i^a) 



Again, simultaneous short-wave and far-red illuminations permit 

 a balancing of rates, and equations 5 and 6 give the rate p^p 

 for less and more than balancing short-wave illumination. Then, 

 from equations 3, ka., 5, and 6 expressions for E and D are 

 derived 



E = l+(2b-l)i-L/ai2 = l+(2b-l)p^/(l-b)p2 (7a) 



D = (2b-l)i = (2b-l)p^/(l-b) (8a) 



E = (l/a)-l (9a) 



