20 



RESEARCHES ON FUNGI 



make an important contribution to the efficient organisation of the 

 hymenium. 



The Cheilocystidia.— Along the free margin of each gill (Fig. 11, 

 a and 6) there is a fringe of irregularly clavate sterile cells which, 

 using the terminology of Topin^ and Knoll, ^ might be called 

 cystidiform cells. However, for reasons which will be more fully 

 explained in connection with a description of Psathyrella dissemi- 



FiG. 11. — Lepiota cepaestipes. Cystidia on the gill-edge, 

 o, transverse section through the edge of a young 

 gill with the hymenium developing. The larger 

 cells below are cystidia. b, a gill-edge seen from one 

 side of the gill and showing the projecting cystidia. 

 c — I, isolated cystidia of different forms ; c and / 

 are unicornuate ; d and e bicornuate ; d, f, h, i 

 and j show contents composed of particles which 

 exhibited Brownian movement. Magnification, 350. 



nata, I prefer to call them cheilocystidia, i.e. cystidia produced on 

 the free edge of a gill. The cheilocystidia of Lepiota cepaestipes have 

 free ends which may be rounded, pointed, or extended into one or 

 two short horns. The horns may be either smooth or coated with 

 small particles (Fig. 11, c to ^). The cheilocystidia are by no means 

 of uniform size, for their length varies from 40 to 70 /j, and their 

 width from 10 to 20 yu,. Each contains a large vacuole in whjch is 



^ J. Topin, Notes sur les Cristaux et Concretions des Hymenomycetes et siir le 

 Role Physiologique des Cystides, These, St. Germain-en-Laye, 1901, p. 29. 



^ F. Knoll, " Untersuchungen iiher den Bau und die Function der Cystiden und 

 verwandter Organe," Jahrb. f. wiss. Bot., Bd. 50, 1912, p. 453. 



