52 RESEARCHES ON FUNGI 



upright positions. They may also assist the basidia by supplying 

 them with water and possibly also with food-stuffs. The sub- 

 hymenium consists of rounded or oval cells about two layers thick, 

 whilst the trama is composed of elongated swollen oval cells with 

 long axes directed perpendicularly to the surface of the pileus. 



Previous authors have overlooked the polymorphism of the 

 basidia which has just been described. Patouillard merely remarks 

 that the basidia are protuberant ; but, as we have seen, this is 

 only true of some of them and not of all. His illustration of the 

 hymenium shows a single protuberant basidium surrounded by 

 paraphyses.i Boudier, in his Icones, has a sketch of a cross-section 

 of the hymenium in which are represented five paraphyses and two 

 highly protuberant basidia.^ Doubtless, like Patouillard, he only 

 noticed the most prominent basidia and failed to observe that the 

 basidia, taken all together, have four different lengths. 



Just as in Lepiota cepaestipes, there are no cystidia on the sides 

 of the gills. Cystidia are also absent from the free gill-edges except 

 for a very short distance from the periphery of the pileus. The 

 hairs, which have already been noticed as occurring on the top of 

 the pileus, are present at the rim of the pileus and appear to pass 

 down below the pileus so as to make a slight invasion of the gill- 

 margins. The best way to detect these marginal hairs is to take a 

 whole pileus from its stipe, invert it, and examine it with the low 

 power of the microscope. It will then be seen that there are basidia 

 on the gill-edges but that there are a few hairs also present on these 

 edges near the pileus-rim (Fig. 27, E, p. 46). These hairs resemble 

 in structure those on the pileus and on the stipe, i.e. they are uni- 

 cellular and have swollen bases and long cylindrical shafts. 



Pilocystidia, Pleurocystidia, Cheilocystidia, and Caulocystidia. 

 — It seems to me that there is room for reform in the nomenclature 

 of those hair-like cells which so often occur on the sides of the gills, 

 on the free gill-margins, on the top of the pileus, and sometimes also 

 on the stipes of many Agaricineae. We might extend the meaning 



^ N. Patouillard, Tabulae analyticae fungorum, Paris, 1883-1886, No. 351, 

 p. 160. 



° E. Boudier, Icones Mycologicae ou Iconographie des Champignons de France 

 principal ement Discomycetes, Paris, 1905-1910, Tome i, No. 85, PI. 140. 



