28o 



RESEARCHES ON FUNGI 



The complete diploidisation of any multicellular and multi- 

 nuclear hymenomycetous haploid mycelium through the agency of 

 a single nucleus or relatively few nuclei derived from a haploid 

 mycelium of opj^osite sex or from a diploid mycelium containing 



cL e 



C I 



D 



-9 



C e I 9 



© 



KiG 



H) 



, 148. — Coprinus lagopus. Diagram to show the diploidisation of a liaploid 

 myceUum (ab) by a diploid mycelium (AB)-{-{ah). No. 1 : the cell / of the 

 diploid hypha f g is growing toward, and soon will meet, and fuse with, the 

 cell (/ of the haploid hypha c d e. No. 2 : fusion has taken place and the (AB) 

 nucleus of / is dividing. No 3 : one of the daughter (AB) nuclei has passed 

 into the cell d, thus diploidising it. No. 4 : a wall now separates the cells 



d and /, and the (AB) nucleus in the cell d is dividing. No. 



i) : 



the wall 

 between the cells c and d is partly broken down, and one of the daughter (AB) 

 nuclei has passed through it from d to c. No. 6 : the (AB) nucleus of the 

 cell d has again divided and has sent one of its daughter (AB) nuclei into the 

 cell c. Thus the diploid cell / diploidised the haploid cell d and the diploid 

 cell d in its turn diploidised first the haploid cell c and then the haploid cell e. 

 Highly magnified. 



nuclei of opposite sex is absolutely dependent on the formation of 

 conjugate nuclei. 



The organisation of the nuclei in conjugate pairs {n)^{n) instead 

 of as isolated nuclei (2w) in the diploid mycelium and fruit-body of 

 the Hymenomycetes results, it is true, in delaying the fusion of 

 nuclei of opposite sex until the basidia come into existence, but it 

 has the great advantage that, iri the diploid tnycelium, each member 

 of a pair of conjugate nuclei retains its identity, so that one member 

 of a 2mir can divide independently of the other member of the pair 



