THE TRANSLOCATION OF PROTOPLASM 129 



scattered on the walls of vacuoles distant from the septa (Fig. 63, 

 B, above the fork in a a, between the septa g and j, and between the 

 septa h and i), they tend to congregate on the walls of vacuoles 

 adjacent to the septa (Figs. 63, B, 64, and 67). 



When streaming is active in a hypha (Fig. 63, B) the Woronin 

 bodies usually remain more or less fixed in position in the walls of 

 the vacuoles, and I have never yet seen them leave those walls, 

 join the granular protoplasm, and be carried off by the current. 

 When they move, they do so spasmodically and their direction of 

 motion is along the wall of a vacuole. If one of them is on the 

 wall of a vacuole occupying the corner of a cell (cf. Fig. 67), its 

 movements along the vacuolar wall may cause it to approach and 

 come up to the septum or to leave the septum. Whilst the majority 

 of Woronin bodies at a septum have their long axes parallel to the 

 septum (Fig. 63, B, g and h, and Fig. 64), some have their long axes 

 more or less perpendicular to the septum (Fig. 63, B, / and j, and 

 Fig. 67). These relative positions are determined by the shape of 

 the vacuole. If a Woronin body moves along a vacuolar wall 

 bulging into the cell-lumen and comes up to a septum, on arrival 

 its axis may be perpendicular to the septum, as shown at /in Fig. 63, 

 B, and to the right in Fig. 67. If then it continues to move and 

 passes on to a vacuolar wall lying against the septum, its axis 

 becomes parallel to the septum. A Woronin body at one moment 

 may be seen in side view as a short rod (Fig. 67) and then it may 

 move round through a right angle and be seen in end view as a 

 rounded structure. An attempt to show Woronin bodies as solid 

 bodies in perspective has been made in Fig. 64 (p. 118). Woronin 

 bodies are less than 1 ^ in diameter and are therefore quite small 

 enough to pass through the pore of a septum. Perhaps they do 

 actually pass through pores occasionally. I once observed a Woronin 

 body on one side of a septum near a pore. After some time, it was 

 found to have disappeared from that position ; and in a vacuolar 

 wall on the other side of the septum in the next cell there were now 

 three Woronin bodies, instead of two as seen previously. The 

 inference is that the Woronin body under discussion had in some way 

 passed through the pore. 



It is not to be supposed that Woronin bodies have any powers 



VOL. V. K 



