i 94 RESEARCHES ON FUNGI 



into the spore, while the other passes downwards into the parent 

 cell (h-o). A spore, therefore, contains but a single nucleus. 



When an old spore is forming a new spore on a sterigma, just as 

 when a yeast cell is forming a spore, the young spore is devoid of a 

 nucleus (Fig. 97, b-d) and, as soon as the new spore has attained 

 almost full size, the nucleus of the parent spore passes into the 

 sterigma and there divides (e-/). Of the two daughter nuclei one 

 passes through the neck of the sterigma into the new spore, while 

 the other passes downwards into the old spore (g-l). Again the 

 newly formed spore contains but a single nucleus. 



A number of recently discharged spores, each containing a single 

 nucleus, are shown in Fig. 98, A. The nuclear phenomena attending 

 the germination of a spore and the production of a bud are similar 

 to those already described for the development of a bud from an 

 ordinary yeast cell, i.e. the young bud is at first devoid of a nucleus 

 (Fig. 98, B, a), the nucleus of the parent spore then passes to the 

 junction of the spore and the bud and there divides (B, b, c) and, 

 finally, one of the daughter nuclei passes into the centre of the bud 

 while the other passes back into the main body of the parent 

 spore (B, d-f). 



The nuclear condition of certain abnormally developed yeast 

 cells and spores is shown in Fig. 99. The cells b, c, and d in A have 

 very long sterigmata but, as usual, there is one nucleus in the spore 

 and one in the parent yeast cell. The cell e has a sterigma with 

 three teeth or branches. Probably a spore was produced on each 

 of the two lower teeth. The spore now on the third tooth, like the 

 parent cell, has the normal single nucleus. In Fig. 99, B, are some 

 spores producing spores. The old spore 6 has two sterigmata. 

 From one sterigma a spore has doubtless been discharged. The 

 spore borne on the other sterigma, like the parent spore, as usual 

 contains but a single nucleus. The old spore c bears three sterig- 

 mata and contains a single nucleus. Probably the sterigmata were 

 developed in succession and three spores were produced and dis- 

 charged in succession. If so, the nucleus of the old spore underwent 

 three successive divisions . At d is shown an old spore with a branched 

 sterigma. Probably the sterigma was at first simple and produced 

 and discharged a spore from the tooth now sporeless, and the 



