GOSSYPIUM 419 



or short hairs. When the lint and fuzz are completely removed, the 

 naked seed is black or dark brown. In some varieties, no fuzz is 

 produced and the seed is left naked after ginning. In Upland 

 cotton, the two kinds of hairs are mixed together over the whole 

 surface of the seed; but, in some varieties, the lint tends to have 

 longer fiber at the rounded end than at the pointed one. The 

 lint is usually white while the fuzz is frequently greenish, brown- 

 ish, or tawny. 



The seed coats are developed from two integuments and have 

 been described in detail by Winton (36). The outer epidermis, 

 aside from the fiber cells or hairs, consists of cells which are 

 irregular in shape. In surface view, they are somewhat stratified, 

 with yellow walls and dark brown contents. They form rosette- 

 like groups around the hairs and vary in size from 10 to 60 ix. 

 Stomata with thin, translucent-walled guard cells occur either 

 singly or in pairs. The hypodermal layer is two to three cells 

 in thickness, except in the region of the raphe where it is thicker, 

 and consists of thin-walled cells with irregular contour and brown 

 pigmentation that may become more or less compressed or crushed. 

 Inside this zone is the innermost layer of the outer integument 

 comprised of rather thick-walled, colorless cells. (Fig. xi}.) 



The palisade cells constitute approximately one-half of the 

 thickness of the seed coat. These elongated cells are very char- 

 acteristic in structure, consisting of an outer portion occupying 

 about one-third of the length of the cell with nearly colorless 

 walls, and an inner portion with yellowish-brown walls. In 

 the outer portion of each cell, the lumen is very narrow except 

 for a globular enlargement at the inner end which contains a dark- 

 colored substance. The inner portion appears to have no lumen 

 when viewed in cross section; and, in the tangential view, radiat- 

 ing lines may be seen which probably result from the arrangement 

 of the lamellae of the cell wall. The inner brown coat consists of 

 several compressed layers of more or less spongy polygonal cells 

 which contain a brown coloring matter such as is found in the outer 

 brown coat. 



Within the inner brown coat is a delicate white sheath two 

 cell-layers in thickness which encloses the embryo. The cells of 

 the outer layer were designated as "fringe cells" by Hanausek 

 (xo), who regarded them as the remains of the perisperm or nucel- 

 lus, and a similar interpretation was made by Winton (36) and 



