258 Report of the Department of Botany of the 



described by the writer (1909) as one new to science. It was found 

 and described only in its imperfect stage and was called Fusicoccum 

 viiicolum. Since that time Shear (1911) has found the ascosporic 

 or so-called perfect stage. He finds it to be one of the genus Crypto- 

 sporella and calls it C. viticola. 



LIFE HISTORY OF FUNGUS. 



Perithecia. — The Cryptosporella stage of the fungus has been 

 reported by Shear from Michigan, Virginia, and New Jersey. This 

 author states further, p. 118, that it is doubtful whether the ascosporic 

 stage plays any important part in perpetuating the fungus. The 

 writer had made diligent search for it in many of the vineyards of 

 the State, and had attempted to develop it in culture, but up to the 

 present time has not succeeded in finding it. 



The perithecia of the fungus, as described by Shear, are buried 

 in a stroma of fungous and host tissue occurring beneath the cortex. 

 The cortex is lifted up but the stroma does not become exposed as 

 in the case of the pycnidia. The perithecium is globose, thin-walled, 

 and provided with a short, smooth beak which extends through 

 and slightly beyond the cortex (Fig. 8). Within the perithecium 



are numerous cylindrical asci measuring 60 

 to 72 by 7 to 8 //, each containing eight 

 ascospores. The ascospores are subelliptical, 

 colorless, one-celled and measure 11 to 15 

 by 4 to 6 n. Between the asci are slender, 

 F IG g_ — Perithecial septate, sterile threads known as paraphyses. 

 Stromata of c. viticrta. The fact that this stage of the fungus has 



never been found in the State although 

 diligent search has been made for it, both before and since the 

 appearance of Dr. Shear's paper, makes it reasonably certain that 

 the perithecial stage of the fungus plays no important role in the 

 dissemination or perpetuation of the fungus with us. 



Pycnidia. — This stage of the fungus is very abundant. It com- 

 monly occurs in the bark of the previous season or of two seasons 

 previous. Fig. 9 (Plate VIII). The young pycnidium forms imme- 

 diately underneath the layer of cork of the last annual ring of bark. 

 Fig. 12 (Plate IX). As it increases in size it lifts up this cork layer 

 and the larger portion of the pycnidium is to be found on the outside. 

 On the berry the stroma develops in the hypodermal parenchyma 



