28 Bulletin 392 



azaleas, sweet williams, stocks, and carnations (Kccble and Armstrong, 

 1912:292). According to De Vries (1905:311, 322, 323), they occur also in 

 liverleaf (Hepatica), dame's violet (Hesperis), larkspur, periwinkle {Vinca 

 minor), tulips, hyacinths, cyclamen, camellia, cockscomb {Celosia crisiata), 

 and Clarkia pulchella, and even in such garden plants as the meadow 

 cranesbill {Geranium pratense). He says (page 323 of reference cited): 

 "It is always the red or blue color which occurs in stripes, the underlying 

 ground being white or yellow, according to the presence or absence of 

 the yellow in the original color mixture." He states further (page 325 

 of the same reference): " Stripes are by no means limited to flowers. 

 They may affect the whole foliage, or the fruits and the seeds, and even 

 the roots." 



In Primula sinensis, altho the amount of flaking varies considerably, 

 the races breed well-nigh true to this habit (Keeble and Armstrong, 

 1912 : 292). 



SOURCE OF MATERIAL 



The source of this character, in the writer's cultures, was in three vines 

 from the stock ntmibered 19, the seed of which was obtained from Dreer's 

 nursery. The seed packet was labeled Dark Blue, but of the six plants 

 that came to bloom from the seeds, one was plain white, one was solid 

 magenta, and four were cream white with magenta flakes. Three of the 

 last-mentioned were selfed and also used in crosses, and their progeny 

 is discussed later. 



THEORETICAL ANALYSIS OF DATA 



The progeny from the three plants mentioned in the preceding para- 

 graph, when selfed, were too few to demonstrate the zygotic constitution 

 of the parents, but indicated that at least two of them were heterozygous 

 in regard to the character of flaking. 



Cream white flaked progeny in the second inbred generation from these 

 three plants were used as parents in 20 crosses in which the other parent 

 was solid-colored. The Fi offspring were dominated by the character of 

 flaking when the flaked parent was (presumably) homozygous for the 

 character. When it was (presumably) heterozygous for the character, 

 part of the offspring were flaked and part lacked the flaking. The 

 Fi flowers were always colored, and where flaking was manifested it ap- 

 peared as a pattern of deeper hue on the solid color. It appeared on 

 purple, dark blue, light blue, magenta, and tinged white corollas. There 

 were 57 individuals in the Fi generation, of which 9 were not flaked. All 

 the remainder showed flaking very strongly except one anomalous indi- 



144 



