I. Inheritance in Shirley Poppij 77 



of cach array of offspring was fouiul in like manner, and tho pink ränge eliminate 

 between this and thc racial Standard deviation. There resulted 



2/(r=-9111, 



or Parcntal Correlation in Colour = ■4122. 



This result, — in excellent accordancc with those for forearm and ^pan in 

 8 cases of 1000 each in man — snggcsts that colour, even with the rough Classifica- 

 tion indicated, may be effectively dealt with. The continuity of the intensity of 

 colouring in Shirley Popj^ies is fairly obviou.s when crops of thousands of flowors 

 arc examincd. We propose next year to fonn a more elaborate scale of colouring 

 and continue the experiments cspecially on the colour side. 



Meanwhile the only comparable data we can use occur in thc Oxford crop. 

 Here the colours were observed on the first flowers of 319 plants distributed 

 among 24 parentages. Owing however to a different colour Classification to 

 W. R. Macdonell's, only 11 of these parentages can be used, — the colour scale not 

 extending over three classes in the others. The mean Standard deviation of these 

 11 parentages — containing only 153 flowers, however, — bears the ratio of '8479 to 

 the Standard deviation of the whole set. Hence we find parental correlation 

 •5302. This result is not very reliable, of course, the data being so few ; yet it 

 is in accordauce with the higher parental inheritance values obtained generally for 

 the Oxford crop. 



(11) Fraternal Correlation. While for resomblance of offspring to parent we 

 have compared an array of offspring plants with the mother plant the seed being 

 taken from one or all capsules of thc mother plant, we have a less definite 

 conception in the case of an array of brother plants. All members of the array 

 have, of course, a common mother plant, they are co-ovarial or bi-ovarial brethren, 

 but the proportions of synanthic, dianthic and dysanthic crossing are quite 

 unknown. Hence it is difficult to guess a priori at the amount of common pollen 

 parentage in pairs of brother plants. We can hardly, however, suppose that all 

 the seed in the sarae capsule has the same pollen parentage, Jmt, except in the 

 case of self-fertilisation, we should expect seed from the same capsule to give 

 plants more alike than seed from different capsules. Taking the best results for 

 fraternities of common parentage in man — mean of about 30 results — we should 

 expect the resemblance of brothers to be about -5. This might be reduced to 

 anything down to '25, i.e. case of half-brothers, by the degree of diversity in the 

 pollen, from which the arrays of brothers are due. Hence we must be prepared 

 for fraternal correlation in poppies lying between •25 and •5, approaching the lower 

 limit, if there has been the freest possible cross-fertilisation, i.e. a great variety of 

 pollen carried not only to the same mother plant, but even to iudividual ovaries 

 on that plant. 



Before we consider actual numerical results we must, however, modify them 

 for homotyposis, remembering that our tables have been formed by comparing 



